Lecture 4: Invertebrates II and Vertebrates Flashcards
Protostomes
- Tripoblasts
- Blastopore becomes mouth first
- Bilateral symmetry
2 Major derived traits:
- Anterior Brain
- Ventral nervous system consisting of longitudinal nerve cords
2 Major clades:
- lophotrochozoans
- ecdysozoans
Lophotrochozoans
What animals?
- Protostomes
- Flatworms. annelids, mollusks
Defining Features of Lophotrochosomes
(not universal)
*likely traits both in a common ancestor, then lost over time
- Lophophore
- circular or u-shaped ring of tentales around the mouth
- food collection and gas exchange - Trochophores
- Free-living larva
- Move by beating a band of cilia
- cilia also bring plankton closer to the larva so it can be captured and ingested
Flatworms
(platyhelminthes)
- Acoelomate
- lack a gas transport system (flat so cells near surface obtain oxygen, no heart or circulatory system)
- move using cilia
- blind gut (only one opening)
- most are parasites (can absorb digested food from host, do not require elaborate feeding or digestive organs)
- sephalization (head with basic chemoreceptors, two simple eyes, tiny brain with longitudinal nerve cords)
- include tapeworms and flukes
Flukes
(flatworm)
Schistosomiasis – snail host
- deposit eggs in open water
- fluke burrows into skin and matures
- migrates to preferred tissues
Tapeworms
(flatworm)
- flea host
- flea is eaten or fecal contamination (meat, water)
- eggs in feces
Annelids
main distinction
(ring worms)
- segmented bodies
- body divided into repeated segments
- ability to move different parts of its body independently of one another
- segments develop from growth zone, oldest in front, youngest in back
ex: bristleworms, earthworms, leeches
Annelids
body systems
- complete gut
- coelomates
- move using muscle contraction
- NERVOUS SYSTEM
- small brain
- ventral nerve cord
- seperate nerve center (ganglion) controls each segment connected and coordinated by ventral nerve cords
- CIRCULATORY SYSTEM
- blood circulates within vessels
- ring vessels near anterior have muscles that act as hearts
- GAS EXCHANGE
- thin permeable body wall serves as a surface for gas exchange
- no gills or lungs
- restricted to moist environments
“Many hairs” Annelids
- bristleworms
- PARAPODIA - thin outgrowths from body wall
- for gasexchange, movements
- covered with stiff bristles for gripping
*paraphyletic - not all in same clade
“pack-saddle” Annelids
- NO parapodia
- hermaphrodites
- thick ring shaped “pack-saddle” around one section of their skin
- holds coccoon in which eggs and sperm are deposited and stored until they hatch
Earthworms - digest soil
leeches - feed on blood, used in medicine tp reduce fluid pressure and prevent blood clotting
Mollusks
main features
- not considered to be truly segmented
- three major body components
- FOOT - muscular structure for support and movement
- VISCERAL MASS - centralized housing for the organs
- MANTLE - tissue covering the visceral mass that can secrete a hard shcll in many species
Mollusks
body systems
-complete gut
- REDUCED coelom (small space around heart)
- instead have a large, fluid filled cavity - HEMOCOEL (cavity for hemolymps - blood and interstitial fluid)
- NERVOUS SYSTEM
- simple brain - nerve ring of connected ganglia
- two pairs of ventral nerve cords
GAS EXCHANGE
- gills
CIRCULATORY SYSTEM
- open system
- heart pumps blood
- blood and other fluids empty into hemocoel
- fluids move around hemocoel and deliver oxygen to internal organs
Types of Mollusks
Chitons
- 8 overlapping dorsal plates
Bivalves
- clams, oysters, scallops, mussels
Gastropods
- snails, slugs, sea slugs
Cephalopods
- squids, octopuses, nautiluses
- enhanced mobility
- modified mantle allos water to be forcibly ejected from the cavity - propulsion
- most complex invertebrate nervous system
- branched foot, tentacles
Ecdysozoans
main types
(protostomes)
- nematodes and arthropods
Ecdysozoans
main features
- defined by having external covering or CUTICLE
- secreted by underlying epidermis
- provides animal with protection and support
- once formed it cannot grow
- must molt and replace cuticle
- can be thin (worm like)
- allows exchange of gases, minerals, water
- restricted animal to moist habitats
- can be thick (arthropods)
- exoskeleton
- impedes movement and passage of oxygen/nutrients
- jointed appendages evolve
- can invade dry land because do not dehydrate
Namatodes (roundworms)
body systems
- Ecdysozoans
- pseudocoelomates
- thick outer cuticle that is shed throughout life
- unsegmented
- complete gut
- GAS EXCHANGE
- through both cuticle and gut
- no gills or lungs
- NO circulatory system
- NERVOUS SYSTEM
- brain = nerve ring
- four peripheral nerves run length of body
How do nematodes eat?
Scavengers:
- soil, bottom of lake or sea
Parasitic:
- multiple stages of life ycle
- heartworms
- hookworms
- pinworms
- whipworms
- trichinella worms
Arthropods
main features
Ecdysozoans
- jointed appendages - complex movement
- most species rich group of animals
- 10^18 alive at any time
- segmented bodes (2 or 3)
- head, (thorax), abdomen
- muscles attached to inside of rigid exoskeleton
Arthropods
body systems
- complete gut
- reduced coelom but have large hemocoel cavity
- open circulatory system
- heart pumps blood through hemocoel cavity
- fluid filled cavity bathes organs in oxygen and nutrients
- no distinction between blood and interstitial fluid: hemolymph
- various systems of gas exchange
- nervous system
- simple brain
- ladder like paired ventral nerve cords with ganglia at each segment
4 Major Arthropod Groups
- Myriapods
- chelicerates
- Crustaceans
- Hexapods
Myriapods
- Millipedes and centipedes
- One (centipedes) or two (millipedes) pairs of legs per segment
- Gas exchange via trachea (no lungs)
*centipedes and millipedes do not have 100 and 1,000 legs
Chelicerates
- Horseshoe crabs
- Arachnids
- Spiders, Scorpions, Mites, Ticks
- Most have 4 pairs of legs
- Many have gas exchange in folded sacs of alternating air & hemolymph
Crustaceans
- Shrimp, Crabs, Crayfish, Barnacles
- Dominant marine arthropod
- Gills which can be located on body or as modified appendages
Hexapods
- Insects
- 1 million species of insects are known.
- Up to 50 million species may exist!
- Dominant terrestrial arthropod
- Six legs on thorax
- Many have two pairs of wings
- Evolutionary advantage
- Unique mechanism of gas exchange
- Sacs & tubular channels that extend from external openings inward to tissues throughout the body
- Complete metamorphosis – changes between molts are dramatic
- Incomplete metamorphosis – changes between molts are gradual
Body cavity in protostomes
- cannot group whole protostomes clade by body cavity type
- likely the common ancestor was coelomate
- loss or partial loss of body cavity seems to have arisen independently several times
Deuterostomes
main features
- blastopore develops into anus
- triploblasts
- coelomates
- no external skeletal suport structures
- some have segmented bodies (vertebrates)
- common ancestors had bilateral symmetry and pharyngeal slits (gills)
- lost by echinoderms
- complex behaviors