Lecture 12 Flashcards
Lignin made of
Complex polymer of phenylpropane units
Lignin Purpose
-Hydrophobic mesh bonds to cellulose and waterproofs the wall,
-Prevents wall enlargement and adds mechanical strength,
-Resists microbial attack,
-It was one of the most important
prerequisites of the evolution of large
upright-growing terrestrial plants,
-Most abundant organic material
on earth after cellulose,
-Lignin stains red with phloroglucinol
Primary vs Secondary Cell Walls
- Secondary walls are less hydrated because they contain fewer pectins and hemicelluloses and because lignin is hydrophobic,
- Secondary walls are less plastic because of the rigidity conferred by lignin,
- Microfibrils in the secondary wall layers are more highly organized and different wall layers usually have a different pitch, which further strengthens the entire wall
- Secondary walls are birefringent (refract light) due to their crystalline arrangement
Primary pit fields become ??? with secondary wall added
Pit Membranes
Bordered pit pairs
Pit membrane with torus in Gymnosperms
Simple pit pairs
No torus
Wood anatomical features observed in RLS
-Bordered pit distribution in axial tracheids,
-Morphology of half-border pit-pairs (between ray parenchyma cells and
axial tracheids),
-Types of rays: Presence of ray tracheids
Sclerenchyma Purpose
Strengthening tissue in a plant, formed from cells with thickened, typically lignified, walls
Sclerenchyma Attributes
- Usually dead cells at maturity,
- With secondary walls containing simple pits,
- Permanently differentiated Found all over the plant body,
- Provide rigid support for mature organs
Collenchyma vs Sclerenchyma
- Collenchyma: Thin primary walls with some areas of secondary thickening. Particularly in regions of new growth,
- Sclerenchyma: Thick lignified secondary walls and often die when mature
Sclereids:
Reduced form of sclerenchyma cells with highly thickened, lignified cellular walls that form small bundles of durable layers of tissue in most plants
Fibers Purpose
Dead at maturity and function as support tissue. Associated with vascular bundles
Fiber Types
Septate, Phloem, Xylem, Gelatinous (little/no Liginifaction)