Lec 11-12 Flashcards
Types of interactions between species
Mutualism: +/+ Commensalism: +/0 Predation: +/- Amensalism: -/0 Neutralism: (0/0) --No interaction b/w species, so not really even an interaction
What do organisms compete for?
Resources:
- Food, water, light (among plants)
- Space
Mates:
- Apparent in animals
- Also in plants, with pollen grains “racing” down the style to the ovary
- Competition within species
Levels of Competition
Intraspecific competition
Interspecific competition
Intraspecific Competition
Competition between individuals of the SAME species
Leads to density-dependent logistic growth curve seen in study of population ecology
Interspecific Competition
Competition between individuals of 2 different species
Exploitation (indirect) competition
Occurs when one competitor uses resources and in so doing reduces the resource availability to the other competitor
-The most common form of competition in all organisms
Interference (direct) competition
Occurs when two competitors physically challenge or harm each other to obtain a resource
- Reasonably common in animals
- Also seen in plants, when one plant releases toxins to harm the competing plant: ALLELOPATHY
Demonstrating competition with Field Experiments: Barnacles
Large acorn barnacles (Semibalanus) and small acorn barnacles (Chthalamus) are both found within intertidal zones
Large acorn barnacles are found throughout the lower and middle intertidal
Small acorn barnacles are found only in the upper intertidal
How can it be shown that this is the result of competition? Do an experiment!
Removal experiment: Barnacles
When large acorn barnacles are experimentally removed, the small acorn barnacles colonize all of the intertidal zone (upper, middle, and lower)
When small acorn barnacles are experimentally removed, the large acorn barnacles remain out of the upper intertidal
Shows:
- Large acorn barnacles are superior competitors, keeping small acorn barnacles out of the lower and middle intertidal
- Large acorn barnacles are susceptible to desiccation, which prevents them from surviving in the upper intertidal
Demonstrating competition with Field Experiments: Paramecium
G.F. Gause experimented with 3 species of Paramecium (unicellular protozoans)
P. aurelia and P. caudatum feed on bacteria
P. bursaria feeds on yeast (unicellular fungus, eukaryotic)
When placed SEPARATE from each other: grow logistically
P. aurelia ad P. caudatum placed together: Both feed on bacteria, P. aurelia is the stronger competitor
P. caudatum and P. bursaria placed together: Coexist; compete for space only, have separate food sources
Competitive Exclusion Principle
Gause established competitive exclusion principle in 1934
Two species cannot coexist in a community if they use the exact same resources in the exact same way
–One species will persist, while the other will be outcompeted and driven to local extinction
Ecological Niche
Niche = When, where, and how a species makes its living
- -All of the environmental factors necessary for a species’ existence (its survival, growth, and reproduction)
- -A species’ role in its environment
- -An n-dimensional hypervolume
- –Abstract, intangible concept
Resource Partitioning
Redefine COMPETITIVE EXCLUSION PRINCIPLE:
-Two species CANNOT occupy the exact same niche within a community
Very similar species can coexist by way of resource partitioning:
-Use the same resources but in slightly different ways to avoid complete competitive overlap
AKA niche differentiation
Modeling Competition
Begin with the logistic equation:
dN/dt = rN (1 - N/K) => dN/dt = rN ((K-N)/K)
Lotka-Voltare Competition Equations:
Species 1: dN1/dt = r1N1 ((K1-N1-aN2)/K1)
Species 2: dN2/dt = r2N2 ((K2-N2-BN1)/K2)
a and B = competition coefficients
a indicates the effect of an individual of species 2 on the individual of species 1
When a=1
If a competitor of species 2 decreases the survival, growth, and reproduction of species 1 by the same amount as another individual of species 1 would, then a=1
Intraspecific and interspecific competition have the SAME effect in this case
When a=2
If a competitor of species 2 decreases the survival, growth, and reproduction of species 1 twice as much as another individual of species 1 would, then a=2
Interspecific competition is then STRONGER than intraspecific competition
Interpretation of B is _________ to interpretation of a
identical
If a and B are both GREATER than 1, the two species ______________-
Could NOT likely coexist
Equilibrium solutions
Set the Lotka-Volterra equations equal to zero to find the population size (N1 or N2) at which the species is in equilibrium (not increasing or decreasing)
Species 1: N1 = K1 - aN2
Species 2: N2 = K2 - BN1
Equilibrium solutions can be demonstrated graphically as ____________________
Zero-growth isoclines
Isoclines show equilibrium sizes for one species in combination with different numbers of individuals of other species
Isoclines can also be used to determine the condition under which each species will increase or decrease in size