FINAL EXAM- Olfaction Flashcards
Is intra- or inter-specific conflict worse? why?
- between group conflicts are worse as escalation occurs faster
- group competition often about territory and resources on those territories
what was the first sensory modality?
olfaction
what other sensory modality is sometimes connected to olfaction?
taste
Flehman response
- licking to bring molecules on tongue directly to the VNO (secondary olfactory system)
- raise head to allow air into mouth and cause molecules to spin, getting them closer to the VNO
what causes the quick processing of molecules during olfaction?
molecules are only one layer away from your brain which allows quick processing compared to other modalities such as vision
what proximity is olfaction designed for?
close proximity–> which causes constraints, and is under environmental constraints (ex. wind direction/ strength)
describe the speed at which volatile molecules travel?
-do not move quickly
=quite stationary
-if no wind, then will stay near source
Name 3 means in which olfactory molecules can be dispersed?
- current flow or fluids (air or water)
- diffusion –> gas expansion
- close to source is highest intensity
- gases expand - direct contact
- gustation often involved- but not actually for taste
implicit processing for olfaction is _____?
=smelling (unconscious)
explicit processing for olfaction is _____?
=sniffing (conscious)
-motor system involved, actively processing info around you
In what 4 dimensions does olfaction differ in?
- Directionality
- high to low concentrations, currents will influence - Speed
- diffusion is slow, trade off for permeance - temporal pattern
- almost none, or lost quickly - Spectrum
- no physical continuous dimensions
what is the production of chemical signals highly dependent on?
- The type of transmission (diffusion, current-borne, contact)
- The medium (air vs. water)
- often species specific
- dependent on density, composition, scent
- liposoluble (more persistent) or hydro-soluble
Contact odours
- usually pheromones
- dependent on what trying to contact
- hydrophobic
- lipids, large hydrocarbons
Waterborne odours
- size restriction of molecules do not count
- large organic compounds
- need to be water soluble (determined by polarity)
Airborne odours
- Volatile: evaporation needed but don’t want too quickly
- Stimulus: molecular size, weight and shape are important
- Environment:
1. wind and convection turbulences
2. Atmospheric conditions (can cause movement)
3. micro-meteorology
4. pedology: soli composition can influence how molecules decoded
describe a scent plume
- if you could see molecules, you would see a scent plume
- related to climatic/ environmental conditions
what are the main interferences with scent plumes?
- wind
- temperature
- humidity
- barometric pressure
what tool can be used to visualize wind?
=smoke bombs
**cant visualize movement of scent molecules just the wind, though scent molecules follow similar rules
what is key to understanding scent plumes?
=stability of the atmosphere
compare surface temp with temp aloft, the greater the difference= the more unstable/ greater the turbulence
what conditions are bad for sniffing?
- cold and dry conditions (i.e. winter)
- high positive ion concentrations
describe ‘tracking’
when an animal has acquired a scent/ trail, and is right on it
head drop= acquired scent
describe ‘trailing’
- before having acquired a source of an odour, looking for it
- related to ‘air scenting’
Plume Pattern: Looping
-high degree of convectional turbulence (diff in temp of ground and higher/ certain altitude)
- daytime
- cool air at high altitude, warm air at surface
-directionality hard to figure out (plume goes up and down, up and down)
Plume Pattern: Coning
- Normal/ stable thermal conditions
- cloudy days
- no dramatic cooling or warming
- great air scenting conditions
Plume Pattern: Fanning
- scent plume compressed vertically and fans out horizontally (higher than ground)
- calm clear night
Plume Pattern: fumigating
- stable air aloft, unstable air on ground (cooler scent plum diffuses down through warmer air)
- sunrise/ dawn
Plume pattern: lofting
- dusk/ sundown
- ground cooling but air still warm
diffusion vs. current flow–> which is dominant
current flow is dominant
What 3 factors influence the gradient of odour transmission?
- velocity (speed)
- viscosity of medium (state of semifluid)
- spatial scale (the ‘space’)
- some animals may have mental representation of space/ plum (at least innate)
Describe Reynolds number (Re)= gradient
=inertial forces/ viscous forces= how likely to stick somewhere
- mean flow velocity, length (distance), kinematic fluid viscosity
- goal is to quantify the transition between laminar and turbulent patterns of flow
explain the results of Reynolds number
small Re= mostly diffusion
moderate Re= laminar flow
high Re= turbulent flow
what is the diffusion coefficient
=D= degrees of freedom of a molecule in the medium
Factors defining D (3)
- size of molecule
- interaction of molecule with medium
- interaction of molecules with each other
what should be known in order to describe a ‘stimulus field’ ?
- the amount of chemical released
2. the threshold of detection for recipient/ reciever
Once a stimulus field is described, what can be determined?
- spreading
- fading
- density
Diffusion example: single, discrete puff
Q= number of molecules released k= receiver detection threshold
Active space:
- Sphere- above ground
- hemisphere= ground level
-relationship between these determines fade-out time
Diffusion example: single, continuous emission
Q= molecules/ sec –> smooth concentration gradient
not efficient way to communicate
Diffusion example: continuous emissions from a moving source
concept of a trail/ or tracking applies here
=sequence of puffs
Ex. snails, slugs, ants, snakes, etc.
Q= number of molecules released k= receiver detection threshold D= degrees of freedom of a molecule in medium L= length of a trail
current flow will increase what two things?
- speed of transmission
2. distance of transmission
Name 2 factors in current flow
- minimal: convection (vertical displacement)
2. maximum: advection (horizontal displacement)
Describe laminar flow
-displacement of molecule unidirectional, without turbulence, with constant velocity
- more likely with:
- slow velocity
- less dense media
- short distances - spread prediction easy, predictable, therefore navigation easier to track plum
- not common in normal environmental conditions (b/c wind, temp differences)
describe turbulent flow
- opposite of laminar: vortices, eddies, waves, etc.
- more likely with:
- high velocity (reduce range of transmission)
- dense media
- long distances
-
flows are created by ___ and ___?
- temperature differential
- Coriolis forces (from rotation of planet)
* cause convention effects
Name the 3 main factors that determine laminar vs. turbulent flow
- velocity (speed) of the flow
- density of medium (water vs. air)
- distance of defined flow space (things fade with distance)
what are hyperbaric conditions?
=high pressure conditions
scent dissipates quickly, but hugs the ground (pressure on ground)
in humans, threshold scores are higher in hyperbaric conditions
what are hypobaric conditions?
=low pressure
molecules NOT compressed to ground, can move freely–> initial scent will travel farther and higher
how does temperature influence taste and smell?
high temp warms things, which increases taste and smell- evaporation occurs faster
what are two interacting factors in terms of temperature?
evaporation and boiling point
how does temp influence evaporation?
- low evaporation at low temp; slow fading
- high evaporation at high temp; quick fading
how does molecular weight impact evaporation?
low evaporation at high molecular weight
at what boiling point are you more likely to smell?
low boiling point
how does rain impact evaporation?
rain keeps odour in ground (not all evaporated) –> if warm day
how does temperature impact turbulence?
high temp= thermal convection= high turbulence
impact of air humidity on spread of odours?
high humidity in air will decrease spread of odours= bad for trailing
humid air is heavier
unless lipid based, will diminish quickly
impact of ground humidity on spread of odours?
high humidity in ground will trap odours at ground level= good for tracking
definition of odorants/ odours/ scent/ smells
generic and/or neutral stimuli
may be organic, but not necessarily with biological significance
definition of ‘semiochemical’
=chemical volatile used in communication, also potentially signification or representation (can communicate a lot of info)
biological significance
definition of ‘pheromones’
for intra-specific communication
‘social odours’
definition of ‘allelochemicals’
inter-specific communication (between species)
kairomones
=subtype of allelochemicals where RECIEVER benefits
allomones
=subtype of allelochemicals where SENDER benefits
synomones
=subtype of allelochemicals
=mutualistic in nature (sender and receiver benefit)–> bidirectional
‘primer effect’ of pheromones
=changes via endocrine and neuroendocrine system (eg. McClintock effect)
prime endocrine system–> any changes will be detectable