final Flashcards
allopatric speciation - vicariance
caused by geographical separation
- ex: oxbow lake formation
- tectonic plates & penguins
allopatric speciation - founder effect
a small subset of indiv separates from original pop & branches into new species
- ex: european starlings
- snails w/ opposite handedness
parapatric speciation
gradient leads to speciation
* birds around tibetan plateau
sympatric speciation
separate areas of same habitat
* sickleback fish
* hawthorn/apple inesects
* stick bugs on adjacent bushes
What is polyploidy? How does it influence reproductive isolation and speciation?
polyploidy is having multiple sets of DNA, which arise from non-disjunction ➞ when indiv mate they produce offspring with odd numbers of chrom who are inviable or infertile
evolutionary radiation
when rapid speciation results in a burst of new species from a single lineage
adaptive radiation
burst of speciation occurs b/c a group of species adapts to new ecological niches
* european finch colonized hawaii & adapted beaks based on their island specific envir
* California tarweed able to adapt to abiotic niches: elevation & precipitation ➞ unfilled space w/ little competition
relative neighbor effect
difference in interactions with neighboring species from low to high elevation
* benefits at ↑ elevation ➞ protection from wind, snow burial, cold sun, UV
intermediate disturbance hypothesis (IDH)
species diversity is highest at intermediate levels of disturbance because competition reduces diversity at low levels of disturbance and death reduces diversity at high levels of disturbance
IDH at low freq mild disturbance
more competitive exclusion ➞ org best suited to that situation/better competitors are ones surviving
IDH high freq, intense disturbance
death
1° succession
bare rock, no soil
* takes very long time to colonize
climax community
successional timeline has completed ➞ stable
2° succession
major disturbance kills plant community but soil remains
* soil ➞ nutrients & anchor for plants
* more rapidly: colonizers can start immediately no need for soil profile to develop
major disturbance events leading to 1° succession
- meteor
- glacier retreat
major disturbance events leading to 2° succession
fire
geographical isolation
prezygotic barrier where org are physically separated and cannot come into contact
habitat isolation
prezygotic barrier of georaphical isolation where org occupy diff parts of same habitat and do not come into contact
* stickelback fish
* stickbug on adjacent plants
* hawthorne insects
mechanical isolation examples
- flower shape
behavioral examples
- flower shape
fires in Mediterranean ecosystems
- Pine Fire syndromes
- chapparal shrub syndromes
Pine Fire syndromes
mediterranean ecosystem fires:
1. fire tolerators: tolerates fire with goal of surviving
* ex: tall, no branches at bottom, very thick bark, long needles
2. fire embracers: lean into fire to trigger next generation
* short, thin, flammable, light up quickly
* open cones & disperse seeds when exposed to heat
* cannoy reproduce w/out fire
chapparal shrub syndromes
mediterranean ecosystem fires:
1. fire recruiters: adult plant dies but has been dropping seeds into soil that are triggered by heat & germinate immediate after fire
2. fire persisters: above-ground portion is burned away by root mass survives & plant can resprout from root mass
prairie fires
intentionally set to promote regrowth
* commensalist interaction btwn trees/shrubs
early successional species characteristics
- short lifespan
- rapid reproduction
- many small seeds
- early reproductive age
- small bodies
- rapid growth
- bad competitors
- boom/bust population growth
- r-selected
late successional species characteristics
- long lifespan
- slow growth
- produce few large offspring
- late reproductive cycle
- better competitors
- large bodies
- stable pop size
- K-selected
competitive exclusion principle
2 species competing for the same limiting resources cannot coexist ➞ eventually the stronger competitor will drive the weaker competitor extinct
ways to avoid competitive exclusion principle
- resource partitioning
- separating habitat into physical parts, like lizards in their parts of tree/shrubs
- separating habitat into “parts” like wavelengths for understory plants or pollinators based on color or shape
- character displacement: species competing for same limiting resources diverge in morphology due to NS
lotka-volterra equation for prey pop
lotka-volterra equation for predator pop
lotka-volterra equation terms that represent reciprocal density dependence in population sizes of predators and prey
V & P
lotka-volterra equation term that represents the growth rate of the predator population
cpV
victim pop size × predation rate × conversion efficiency
why are the lotka-volterra equations formulated from the exponential growth equation and not from the logistic growth equation
they believed that the primary driver for pop size for each was the others’ pop size & neither would reach K so it is irrelevant
Why might natural selection favor a predator that is LESS efficient, or a disease/pathogen that is LESS virulent?
- if a predator is too efficient then it drives prey extinct & itself following
- If pathogen kills host then it dies too
consumption efficiency
how much do you eat of the amount of biomass available
assimilation efficiency
how much of what you consume is digested
production efficiency
how much biomass can you produce from what you digest
ecological efficiency
proportion of net primary energy that becomes net secondary energy
consumption x assimilation x production efficiencies
Lindeman’s law of 10%
~10% of energy available at one trophic level is transferred to the next
to determine how many trophic levels an ecosystem can support
- available energy through primary productivity
- efficiency of energy transfer across tropic levels
important roles/identities that org have in their trophic interactions
- keystone species: very high impact on diversity despite being rare
- foundation species: physical bodies make up habitat for other org
- ecosystem engineer species: physically alter habitat
keystone species examples
- prairie dogs burrows house other org & create patchiness gives diversity of plants that attract diverse primary consumers ➞ promotes diversity at all higher trophic levels
- seastars control prey pop sizes which increases diversity by preventing competition btwn prey species
foundation species examples
- corals
- trees
- kelp
ecosystem engineer species
beavers’ damns alter water flow & promote species richness
tropical rainforest
- high precipitation w/ little/no seasonality
- high temp with no seasonality
Desert
- extremely low precip with little/no seasonality
- generally high temp with high seasonality
Temperate deciduous forest
- medium precip (higher than grassland but not as high as rainforest) with no seasonality
- medium temp with seasonality
Grassland
- high seasonality in temp and precip
- wet season occurs during the warm season
Boreal forest
- low precip with some seasonality
- generally low temperatures (~6 months below 0 C) and high seasonality in temp
Tundra
- low precip (but not as low as a desert) with some seasonality
- very low temperatures (~9 months below 0 C) and high seasonality in temp
Mediterranean
- high seasonality in temp and precip
- wet season occurs during cold season
tropical rainforest
desert
temperate deciduous forest
Boreal forest
artic tundra
temperate grassland
mediterranean
Anthropocene
era when human activities are the dominant influence on climate & the envir
grassland in S
tropical rainforest at equator
desert in N
temperate deciduous in N
desert in S
mediterranean in N
grassland in N
mediterranean in N
arctic tundra
boreal forest in N
tropical rainforest at equator
mediterranean in S
Would you expect the production efficiency to be higher for an ectotherm or an endotherm?
ectotherms
aquifers vs oil, & coal
‘fossils’ as they come out of the ground, are slow to replenish but they do not form from fossils
extinction vortex
path to extinction where every step pushes a species closer and closer
1. human or natural disturbance event
2. smaller pop size results
3. genetic drift, inbreeding, random pop size decrease genetic diversity & make pops more susceptible & less likely to withstand
4. reduced fitness
5. lower reproduce & higher mortality
6. smaller pop ➞ cycle continues until extinction
Nₜ = N₀ert
predicting population size under exponential growth
age demographics and r
r correlates to the proportion of indiv in their reproductive ages
- ↑ proportion of indiv in reproductive age = higher growth rate
- ↓ proportion of indiv in reproductive age (majority of pop post-reproductive age) = slower growth rate
both homozygotes are overrepresented
- inbreeding
- natural selection- diversifying
hamilton’s rule
defines how benefits to close relatives (↑ reprod output) can outweigh costs to the altruist (own lost reprod output from altruistic event)
- when is kin selection supported by natural selection
r B > C
r = coefficient of relatedness: fraction of genes shared
B = benefit to relative: ↑ in offspring for relative
C = cost to altruist: loss of offspring for altruist
- ↑ benefits or relatedness incurs a higher cost
exceptions to hamilton’s rule:
- reciprocal altruism: altruist has reasonable expectation that sacrifices will be reciprocated in the future
- repeated interactions
- non-related indiv
- ex: vampire bats
- sexual selection: displays of altruism ↑ mating options
linkage
genes close to each other on the same chrom will be inherited together
- disproportionate ratios btwn gametes
- if AB are on same chrom & BC are on the same chrom, then A & C are on the same chrom
recombination freq
genes on diff chrom: expect nearly equal proportion of the 4 diff types of gametes
genes on same chrom: proportions of gametes are not equal
- majority = non-recombinant (linked)
- minority = recombinant
pleiotropy
one gene affects multiple diff traits/phenotypes
polygenic inhertence
one trait is additively controlled by many genes
- phenotype controlled by combination of many different genes
- every gene is allowed to act
- continuous distribution
- range/variation of phenotypes
- ex: height, color
epistasis
multiple genes interact to determine phenotype
- one gene can mask another
gene flow
migration: movement of alleles through indiv or their gametes
- new allele introduced to pop ➞ changes allele/gene freq ➞ starts out low but allele becomes more common
- w/in pop: ↑ gene variation
- homogenize distant gene pools/pop
- org can migrate without moving
- ex: pollen, marine animals
genetic drift
chance events in small pop cause unpredictable changes in allele freq
- random
- rare alleles are lost entirely after a single generation in small pop
- random event effects are stronger in smaller pop
- graph w/ many squiggles
- reduces gene diversity
- ↑ homozygosity ↓ in heterozygosity
- can be stronger than NS
when given observed & expected w/in pop size, cannot predict which direction drift will shift in freq
consequences of genetic drift:
- loss of overall diversity through loss/fixation of alleles
- no gen diversity ➞ cannot adapt
- ↑ in deleterious recessive conditions
- ↑ susceptibility to future stressors
random mating
only occurs when every indiv has an equally likely chance of mating with another
HWE: non-random mating influences
- sexual selection
- mate preference
- proximity
inbreeding
mating between relatives
- familial relatives
- ex: cousins, 2nd cousins etc
↓ level of heterozygosity & ↑ homozygosity compared to expected
HWE: non-random mating consequences
- inbreeding ↓ heterozygosity & ↑ homozygosity
- homozygotes are overrepresented
- inbreeding ↑ freq of deleterious recessive alleles
HWE: NS consequences
- fit alleles are overrepresented in future gen
- ↓ freq of unfavorable traits
- favors particular alleles over others
- directed selection
Nᵪoff
of offspring of indiv at age x
lᵪ
survivorship: where is mortality primarily occurring
- lᵪ = Nᵪ ÷ N₀
mᵪ
fecundity: avg # of offspring each indiv has at that age
- mᵪ = Nᵪoff ÷ Nᵪ
lᵪmᵪ
what age group has the most offspring
G
generation time: avg age of reproduction
- should be close to age where most reproduction occurs (lᵪmᵪ)
- G = ∑ xlᵪmᵪ ÷ ∑ lᵪmᵪ
R₀
net reproductive rate: avg # of offspring each indiv has throughout their lifetime
- R₀ = ∑ lᵪmᵪ
- R₀ = 1 ➞ pop size is not changing
- cannot be negative
relationship between R₀ and r
when R₀ = 1, r =1 ➞ every indiv is replacing themself in future gen
when R₀ > 1, r > 0
when r < 0, R₀ < 1 (but greater than 0)
what contributes to genetic diversity?
- sexual reproduction mixes whole genomes from 2 diff pop
- independent assortment mixes sets of chrom & allows org to make genetically unique gametes
- recombination mixes alleles on a chrom ➞ creates new chrom combinations diff than inherited
- mutations
one homozygote is overrepresented
long-term drift
heterozygotes are overrepresented
- outbreeding
2 natural selection - stabilizing