Exam 4 Flashcards
animals are ______, ________ eukaryotes with ______ that develop from embryonic ________
multicellular, heterotrophic, tissues, layers
what several characteristics when put together define the animal kingdom
Nutritional mode:
- Heterotrophic. No photosynthesis (not a plant)
- internal digestion (not a fungus)
Cell Structure and specialization:
- multicellular eukaryotes
- cells do not have cell walls
- specialized tissues (esp nervous and muscle) that develop from layers
Reproduction:
- Sexual reproduction
- sperm and eggs directly by meiotic division
- often larval stages
Early embryonic development in animals
- Zygote of an animal undergoes a series of mitotic cell divisions called cleavage
- An 8 cell embryo is formed by three rounds of cell division
- In most animals, cleavage produces a multicellular stage called a blastula.
- Most animals also undergo gastrulation
- the pouch formed by gastrulation (archenteron) opens to the outside via the blastopore
- the endoderm of the archenteron develops into the tissue lining the animals digestive tract
what is a blastula
a hollow ball of cells that surround a cavity called the blastoceol
what is gastrulation
a process where one end of the embryo folds inward, expands, and eventually fills the blastocoel, producing layers of the embryonic tissues; the ectoderm and the endoderm
larva
larva is sexually immature and morphologically and behaviorally distinct from the adult stage. Most animals have at least one larval stage. Larva may even have a different habitat than the adult. They will undergo metamorphosis where they become juveniles that resemble adults but are sexually immature.
what are the developmental genes that regulate the expression of other genes that all animals have called
hox genes
animals are more closely related to what than plants
fungi
morphological and molecular data indicates that protists called what are the closest living relatives to animals
choanoflagellates
multicellularity requires cells to do what
adhere and signal to each other.
animal genes involved in adherence have sequence similarities in choanoflagellates
Timeline of animal development
Neoproterozoic era: - chemical evidence - ediacaran biota Paleozoic Era: - cambrian explosion - anthropods to land - vertebrates to land Mesozoic Era: - age of reptiles Cenozoic Era: - age of mammals (current day)
Neoproterozoic Era
- Chemical evidence of steroids used by sponges has been found in 710 million year old sediments
- Molecular analysis suggests the common ancestor animals likely lived about 770 million years ago
- Ediacaran biota the first macroscopic animals
- Possible predation
Paleozoic Era
- the cambrian explosion marks a period of rapid animal diversification
- most fossils from cambrian explosion are bilaterians (bilateral)
- land colonization: anthropods and vertebrates
Mesozoic era
- coral reefs- huge influence on marine habitat
- reptile diversification. Dinosaurs emerged as dominant predators and herbivores
- mammals (tiny nocturnal insect eaters) appeared
- flowering plants and insects diversified
cenozoic era
- beginning of this era followed mass extinctions of both terrestrial and marine animals
- dinosaurs and marine reptiles extinct
- mammals increased in size and abundance
- the primate diversification
how can animals can be characterized by body plans
- symmetry: 3 types (radial, bilateral, or asymetric)
- Tissues layers (2 or 3)
- Body cavities (3 types)
- protosome and deuterostome development
Body cavities
- most triploblastic animals have a body cavity (a few don’t) a fluid or air filled space between the digestive tract and the outer body wall
- Body cavities have many functions:
- The internal fluid cushions the suspended organs
- The fluid can act like a skeleton against which the muscles of soft bodied animals can work
- The cavity enable internal organs to grow and move independently of the outer body wall
- compact (no body cavity), Hemocoel, or coelom
protostome and deuterostome development
- Bilateral animals can be catergorized as having one of two developmental modes: protostome development or deuterostome development
- these modes differ in cleavage, coelom formation, and fate of the blastopore
Cleavage in protostome development vs. Deuterostome development
Protostome development: - spiral cleavage - determinate cleavage Deuterostome development - Radial cleavage - indeterminate cleavage
radial symmetry
body parts are arranged around a single main axis that passes through the center of the animal. Axis divides the body into mirror images
bilateral symmetry
body parts of the animal are arranged around two axes of orientation, the head-tail axis and the dorsal-ventral axis. Only one imaginary slice divides the animal into two mirror images referred to as the left and right sides
fate of blastopore in protostome vs deutersostome development
- blastopore is an indentation in the gastrula that leads to the formation of the archenteron
- the blastopore and second opening at the opposite end will form the mouth and anus: in protostomes the blastopore becomes the mouth. In dueterostomes the blastopore becomes the anus
what are several data sources used to infer evolutionary relationships among the three dozen extant animal phyla
- morphological traits
- whole genomes
- ribosomal RNA (rRNA) genes
- Hox genes
- protien coding nuclear genes
- mitochondrial genes
the 5 key ideas about phylogeny of living animals
- All animals share a common ancestor: that animals are monophyletic, forming a Metazoa clade, meaning they all have descended from a common ancestor
- Sponges are the sister group to all other animals: sponges are basal animlas that diverged from all other animals early in the history of the group
- Eumetazoa is a clade of animals with tissues:
All animals except for sponges and a few others belong to a clade of eumetazoans. They have tissues such as muscle tissue and nervous tissue - Most animal phyla belong to the clade bilateria:
bilateral symmetry and the presence of three prominent germ layers are shared derived characteristics defining the clade bilateria. - There are 3 major clades of bilaterian animals:
Biliaterians have diversified into 3 main lineages: deuterostomia, lophotrochozoa, and ecdysocoa.
sponges (porifera)
basal animals that lack tissues
do not undergo gasturlation
sedentary and radial symmetry
filter feeders: pores –> spongocoel —> osculum
Flagelluym = flow
choanocytes and amoebocytes
Reproduction: sequential hermaphroditism. Mobile, falgellated larvae
Cnidaria
Eumetazoans, animals with true tissues
gastrula –> diploblastic
Radial symmetry, gastrodermi with single opening
Medusa/polyp sensory structure
muscles
Two major clades: Medusozoa and Anthozoa. Medusozoa forms alternate polyp and medusa. Anthozoa is polyp only
lophotrochozoan
- Flatworms: freeliving, termatodes, tapeworms
- Rotifers
- Ectoprocts & Brachiopods
- Molluscs: gastropods, bivalves, cephalopods
- Annelids
Flatworms
Phylum Platyhelminthes includes flatworms which are dorsoventrally flattened and have no body cavity
Single opening digestive system
free-living and parasitic species
Parasitic flatworms: trematodes, tapeworms
Rotifers
- Flow-through alimentary canal ( finally have an anus which is a big step forward because of flow through digestive system)
- specialized organ systems
- body cavity (hemocoel)
- parthenogenesis is common: asexual reproduction unless they are stressed then sexual reproduction begins
ectoprocts and brachiopods
ectoprods and brachiopods share several traits:
- a lophophore is used for feeding
- a u-shaped alimentary canal
- the absence of a distinct head
- a coelem and deuterstome development
Molluscs
- Phylum Mollusca includes snails and slugs, oysters and clams, and octopuses and squids
- No huge evolutionary step forward, but they are abundant and important in the biotic communities
- mantle, foot, and radula
Gastropods
- about 3/4 of molluscs are gastropods
- single shell
- most are marine, but there are also freshwater and terrestrial species
- gastropods move slowly by using cilia or by a rippling motion of the foot
- feeding: grazers, using radula
Bivalves
- bivalves are all aquatic
- they have shell divided into two hinged halves drawn together by adductor muscles
- they have no distinct head or radula (they are filter feeders)
- some have eyes and sensory tentacles along the edge of their mantle
- bilaterally symmetric; most have sensory organs along the edge of each opening valve
cephalopods
- marine predators
- they immobilize prey using a poison in their saliva
- water is drawn into the mantle cavity and shot out through the excurrent siphon for rapid movement
protecting freshwater and terrestrial molluscs
- molluscs have the largest number of documented extinctions among animlas
- freshwater bivalves and terrestrial gastropods are the most severely threatened
annelids
- segmented worms that live in marine, freshwater, and damp soil habitats
- they have a coelom (no hemoceol)
- based on phylogenomic analysis, annelids are now divided into two clades
Ecdysozoans are the most _____-______ animal group
species-rich
- animals with a cuticle, a tough external coat
- The cuticle is shed during ecdysis, or molting
- primary cavity = hemocoel
- nematodes and arthropods are the largest of eight ecdysozoan phyla
nematodes
- Nematodes (roundworms) have cylindrical bodies tapered at the ends and covered by a cuticle
- They range in length from less than 1 mm to more than 1 m
- Many important animal and plant parasites
- C elegans
Arthropods
- Zoologists estimate that there are about a billion billion arthropods living on earth
- More than 1 million species have been described
- Two of every three known species are arthropods
- Arthropods are found in nearly all habitats on earth
Arthropod origins
- First animals to crawl up on land
- The arthropod body plan = segmented body, hard exoskeleton, and jointed appendages
- They date back to the Cambrian explosion
- Early arthropods ( such as trilobites) showed little variation from segment to segment
- Over time, segments united to form “body regions” specialized for feeding, walking, or swimming
General characteristics of arthropods
- segmented boyd with hard exoskeleton of chitin
- appendages are jointed and come in pairs
- arthropod appendages have become modified for walking, feeding, sensory reception, reproduction, and defense
- open circulatory system
Living arthropods consist of three major lineages that diverged early in the phylums evolution:- Chelicerates (spiders, horseshoe crabs, scorpions, ticks, mites)
- Myriapods (centipedes and millipedes)
- Pancrustaceans (insects and crustaceans)
Chelicerates
- named for clawlike feeding appendage called chelicerae
- arthropod
Myriapods
- Mryiapoda includes millipedes and centipedes
- All living myriapods are terrestrial
- They have a pair of antennae and three pairs of appendages modified as mouth parts
Pancrustaceans
- Recent evidence indicates that insects are more closely related to crustaceans than myriapods
- So, together, insects and crustaceans form the clade Pancrustacea
Crustaceans
- Many crustaceans have highly specialized appendages
- the anterior-most form two pairs of antennae
- three or more pairs are modified as mouth parts
- walking legs are located on the thorax
- smaller, swimming appendages are located on posterior segments (“tail” region)
- Small crustaceans exchange gases through the cuticle; larger crustaceans have gills
- most crustaceans have separate sexes
Isopods
- one of the largest groups of crustaceans, live in terrestrial freshwater, and marine habitats
- Pill bugs are common terrestrial isopods that live under moist logs and leaves