Exam 3 Flashcards
Perceptual process of hearing
1.Sound stimulus is produced
2.Sound travels through the air and is received by the auditory receptors
3.Signals are transduced and sent to the brain
4.Sound information is processed in the brain
5.We perceive the sound
6.We recognize the sound
7.We act on the sound
Physical definition of sound
pressure changes in the air
Perceptual definition of sound
experience we have when we hear
Sound occurs when…
the movement or vibration of an object causes pressure changes in a medium that can transmit vibrations
Condensation
Increase in density
Rarefaction
Decrease in density
Sound wave
pattern of pressure changes
Pure tone
tone with pressure changes that can be described as a single sine wave
Building blocks of sound
Frequency
number of cycles per second that the pressure changes repeat
measured in Hertz (Hz)
Associated with pitch
What range of Hz can humans perceive?
20-20,000 Hz
Amplitude
Size of the pressure change
measured in decibels (dB)
associated with loudness
Decibels increase logarithmically
an increase of 20 dB means the amplitude is 10x greater
Periodic tones
tones with a repeating waveform
Periodic tones have a ____.
Fundamental frequency
Fundamental frequency
Number of times a sound repeats per second
The first harmonic or fundamental of a complex tone, is usually the ___ in the frequency spectrum of a complex tone.
lowest frequency
Higher harmonics
The other components of a tone; frequencies are whole number multiples of the fundamental frequency
Loudness
Perceptual quality most closely related to amplitude of an auditory stimulus
What Hz are humans most sensitive to?
2,000-4,000 Hz
Pitch
quality of a sound ranging from low to high, most closely related to a frequency of a tone
Octave
tones that have frequencies that are binary multiples of each other
Timbre
quality that distinguishes between two tones that sound different even though they have the same loudness, pitch, and duration
Attack
Buildup of sound at the beginning of the tone
Decay
decrease in sound at the end of the tone
Aperiodic sounds
sound waves that do not repeat
Physical qualities of sound
Frequency
amplitude
harmonic structure
Perceptual qualities of a sound
Pitch
loudness
timbre
Three main sections of the ear
Inner ear
middle ear
outer ear
Tympanic membrane
ear drum
What makes up the outer ear?
Pinna
auditory canal
Tympanic membrane
The outer ear is responsible for
resonance
resonance frequency
Resonance
Certain frequencies are enhanced
What makes up the middle ear?
Ossicles
Oval window
middle ear muscles
What are the three parts of the ossicles
Malleus
Incus
stapes
What are the ossicles responsible for
concentrating the vibration of the large tympanic membrane onto the much smaller stapes
middle ear muscles
dampen loud sounds and our own sounds
Parts of the inner ear
Cochlea
cochlear partition- extends from base to apex
organ of Corti
hair cells-stereocilia
basilar membrane
tectorial membrane
Steps of sound transmission
- Stapes vibrates
- Oval window moves back and forth
- Vibrations travel through cochlear fluid
- Basilar membrane moves up and down
5a. Organ of Corti moves up and down
5b. Tectorial membrane move back and forth - Stereocilia of hair cells bend one way
- Tip links stretch
- Tiny ion channels open
- Potassium (K+) flows in
- Electrical signal results
- Neurotransmitters released in synapse
- Stereocilia bend other way
- Ion channels close
Vibrations bend the
stereocilia
bending of stereocilia causes
electrical signals
A sound wave’s frequency determines the ____.
Timing of electrical signals
Phase locking
firing of auditory neurons in synchrony with the phase of an auditory stimulus
Békésy discovered …
how the basilar membrane vibrates like a traveling wave.
Place of greatest vibration depends on…
Frequency
Base (____)
Apex (____)
High frequencies
low frequencies
The basilar membrane has a ___ organization
Tonotopic
Tonotopic map
orderly map of frequencies (tones) along the length of the cochlea
Cochlear amplifier
expansion and contraction of the outer hair cells in response to sound sharpens the movement of the basilar membrane to specific frequencies
Place theory
says that pitch perception is based on the place along the basilar membrane at which the nerve firing is highest
Problem with place theory
amplitude-modulated noise – noise that isn’t associated with vibration of a particular part of basilar membrane, yet still results in pitch perception
Frequency theory
says that pitch perception is based on the frequency of action potentials in auditory nerve neurons, which occur at the same frequency as the sound.
- considered the best theory
Damage to inner hair cells
loss of sensitivity
Damage to outer hair cells
loss of sensitivity and loss of sharp frequency tuning (cochlear amplification)
Auditory pathway
Cochlea
Auditory nerve
Cochlear nucleus
Superior olivary nucleus
Inferior colliculus
Medial geniculate nucleus (thalamus)
A1 (primary receiving area)
Other areas in cortex
Presbycusis
hearing loss caused by hair cell damage resulting from cumulative effects over time
caused by noise exposure, drugs that damage hair cells, and age-related degeneration
affects men more than women
Greatest loss of hearing for Presbycusis
greatest loss for high frequencies
Noise-induced hearing loss
occurs when loud noises cause degeneration of the structures involved in hearing
Noise-induced hearing loss can involve damage to
Organ of Corti
hair cells
auditory nerve fibers
Cochlear implants
use electrodes inserted into the cochlea to create hearing by electronically stimulating auditory nerve fibers.
Parts of cochlear implant
- microphone
- sound processor
- transmitter
- array of electrodes
essentially acts as the hair cells
Auditory localization
perception of the location of a sound source
Auditory space/ scene
the sound environment, which includes the locations and qualities of individual sound sources
Location cues
characteristics of a sound that provide info regarding location of the sound source
Binaural cues
require two ears
determine the azimuth of sounds
monaural cues
requires only one ear
Three dimensions in auditory space
Azimuth
elevation
distance
Azimuth
left-right sound cues
Elevation
up-down sound cues
Distance
how far or close the sound is
Interaural time difference (ITD)
difference between when a sound reaches the left ear and when it reaches the right ear
between-ear sound difference
What is ITD best for
low-frequency sounds
most important binaural cue
Interaural level difference (ILD)
difference in sound pressure level (amplitude of the sound reaching the two ears)
Acoustic shadow
The head blocks the ear, resulting in the sound appearing quieter in the opposite ear
ILD is best for
high-frequency sounds
The ILD and the ITD leave ____ ambiguous
elevation
Cone of confusion
surface in the shape of a cone that extends out from the ear; sounds originating from different locations on this surface all have the same ITD and ILD, so location info provided by these cues is ambiguous
The anterior auditory cortex is important for
pitch perception
A1 travels to other cortical auditory areas
core area
belt area
parabelt area
Core area
A1 and nearby
Belt area
surrounds and receives signal from core
Spectral cue
distribution of frequencies reaching the ear that are associated with specific locations of a sound, caused by interaction of sound with head and pinnae
important for elevation
Parabelt area
receives signals from belt area
Hoffmann et al.
demonstrated the importance of the pinnae for localization of elevation
The Jeffress neural coincidence model
neural circuit for processing the interaural time difference
neurons are wired to each to receive signals from the two ears, so that different neurons fire to different ITDs
Coincidence detectors
neurons detecting the coincidence of both ears firing together
Mammals have much broader
ITD tuning curves