evolution Flashcards
where does trait evolution occur on a phylogenetic tree?
internodes
tips
summations over their evolutionary history
relationship between relatedness and similarity
-similarity does not determine relatedness
-the number of trait changes does not determine relatedness
separate ancestry
the alternative hypothesis to common ancestry, which proposes that each living taxon has an independent origin
rate of trait evolution
-about the same for all branches
-the expected number of changes from the root will be the same for all living tips
-more nodes does not equal more trait changes
-all living species are equally “evolved”
exceptions to the nesting pattern within the common ancestry model
-a trait might evolve independently a few times
-a trait may evolve and later on be lost on one or more lineages
homologous trait
trait shared in separate species that derives from the same evolutionary origin
non-homologous trait
trait shared in separate species that derives from different evolutionary origins through convergent evolution
principle of parsimony
the hypothesis that invokes the fewest changes is most likely to be true; this reasoning works so long as the trait is one that evolves relatively rarely
occam’s razor
similar to parsimony; the simplest explanation for an observation is most likely the correct one
are trait gains or losses more likely?
equally likely
convergent evolution
the same trait evolves separately in more than one lineage; although they appear similar, these traits are not homologous
reversal
an ancestral trait was lost and then re-evolved along a lineage; makes the trait non-homologous
ancestral trait reconstruction
if we know where changes occurred, we also know ancestral states
evolution’s 3 big ideas
1) common ancestry unites all life
-diverse living species descend
from common ancestors
2) populations evolve
-the genetic composition of
populations changes over time
3) natural selection provides direction
-adaptations are explained by
natural selection (and related
processes)
phylogenetic tree
a branching diagram used to represent evolutionary relationships and relatedness between different organisms based on their common ancestry
root
the base of a tree, representing the common ancestral lineage of all taxa in the tree
branch
the lines that make up a tree diagram, which represent population lineages
taxon (plural= taxa)
a named group of biological organisms, often shown at the tips of a tree
lineage splitting/speciation
splitting of a population into genetically separate populations that no longer have gene flow; the origin of multiple species from a few ancestral species
node
branching point on a tree diagram, which represents lineage splitting
reasons for lineage splitting
-geographic changes
-climate changes
-rare dispersal events
clade
all descendants of an ancestral lineage
sister clade/lineage
lineages or clades from the same node
tree topology
a list of all clades a tree contains; stable to pruning
relatedness
recency of common ancestry
biogeography
the geographic distribution of living organisms
common ancestry
the concept that if you trace back the lineages of living species far enough in time, those species will converge to a shared ancestor
evidence for common ancestry
-biogeography
~similar species occur near one
another
-fossil record
~transitional forms in temporal
sequence
-homology
~surprising similarities among
organisms
-classification
~hierarchical nesting of biological
taxa
homology
traits whose similarities are explained by common ancestry
transitional fossil
fossil taxa that have some, but not all, of the derived traits of a living group
nested hierarchy/classification
a pattern of groups nested within groups (without overlaps) as seen in taxonomies
polymorphism
the existence of multiple variants within a population
evolution
the change in frequency of genetic variants within a population; changes in allele frequency of a population over time
fixation
the loss of all variants except one from a population
natural selection
the tendency for genetic variants that enhance fitness to go to fixation and variants that reduce fitness to be lost from a population, making the population better suited to its environment over time
plasticity
changes caused by the environment that do not have a genetic basis
locus
a place in the genome where alleles reside; in diploids, each individual has two alleles per locus
allele frequency
in a population, the proportion of all alleles at a locus that are of a particular type; changes in allele frequency over time causes evolution
heritable variation
allelic variation is segregating in a population
what determines phenotype of an individual?
the alleles they carry at many loci and the environment
hardy-weinberg assumptions
-no mutation
-no migration
-no selection
-mating is random with respect to the alleles
hardy-weinberg law
assuming assumptions hold:
-homozygote frequencies are the
square of the corresponding allele
frequency (p^2 or q^2)
-heterozygote frequencies are twice
the product of their allele
frequencies (2pq)
deviations from hardy-weinberg assumptions that cause evolution
-selection: alleles do not have equal fitness
-genetic drift: population is not infinitely large
probability that an allele becomes fixed under genetic drift
equal to that allele’s frequency
genetic drift
random change in allele frequencies in a population over time