Biology Exam 2: Origins of Life Flashcards
When did the first eukaryotes evolve?
1.5-2 billion years ago
What is used as an evolutionary proxy for early eukaryotes and why?
Living, single celled eukaryotes (protists) are often used as a proxy because the fossil record does not capture the evolution of these single celled euk. as clearly as we would like
monophyletic
sharing a single common ancestor
Prokaryotes v Eukaryotes cytoskeleton? Division? chromosomes? nucleus? organelles? size of cells? number of cells?
Prokaryotes // Eukaryotes
no cytoskeleton // have a cytoskeleton
binary fission // mitosis
circular chromosome // linear chromosome(s)
DNA not enclosed // DNA enclosed in nucleus
No organelles // membrane bound organelles
small // large
typically unicellular (biofilm) // often multicellular
Key evolutionary changes prok –> euk : (6)
- The loss of a firm cell wall
- Origin of a flexable cell surface
- Origin of a nuclear envelope
- Origin of digestive vacuoles
- Origin of mitochondria and chloroplasts by endosymbiosis
- Lateral gene transfer from bacteria and archaea
What had to happen for infolding of the plasma membrane? What did this change allow for (2)?
Loss of a firm cell wall.
- increased cell surface area for higher rates of gas and nutrient exchange
- Endocytosis
Endocytosis played a key role in the evolution of which 3 structures?
the nuclear envelope, membrane bound organelles, and the symbiosis with prokaryotes that produced the chloroplasts and mitochondria
Evolution of mitochondria:
Mitochondria originated as proteobacteria that entered the cell via encapsulation. the proteobacteria developed a symbiotic relationship with the early eukaryote because the mitochondria detoxified free oxygen within the cell
Evolution of chloroplasts:
Chloroplasts evolved via encapsulation of cyanobacteria and the secondarily encapsulated in a larger euk in different lineages. Differing chloroplast membranes in different lineages suggests this happened independently multiple times
The evolution of a cytoskeleton allowed for: (2)
- The physical support of a larger cell
2. The effective transport of material within the cell, including the equal split of DNA into daughter cells
What structure likely evolved from the microtubules found in the cytoskeleton?
flagellum, which allows for proficient propulsion and movement
What are protists or protista?
single celled eukaryotes. They are the most ecologically diverse of all eukaryotes.
Why are protists of great interest to biologists? (2)
- They can be used as an evolutionary proxy to examine the evolution of the first eukaryotes
- Protists are key members of the ecology of our planet, a few are important disease causing agents
Why are protists important in ecosystems?
- Protists include photosynthetic primary producers that are the base of many food chains and produce free oxygen
- many heterotrophic protists serve as a trophic bridge between bacteria and larger organisms
Do protists reproduce sexually or asexually?
Both. Asexual: binary fission, multiple fission, budding, and spores. Sexual: varies. One example is conjugation
Are fungi more closely related to plants or animals?
Animals
T or F: there are still areas of major phylogenetic uncertainty in the evolution of eukaryotes
T
Alveolates
Protists. large, diverse group having sacs (alveoli) under their plasma membrane. Many are typically photosynthetic with cellulose in their cell walls. Contains Apicomplexans (aka sporozoans), Dinoflagellates, and ciliates.
Apicomplexans (sporozoans)
Alveolates. Obligate parasites named for their mass of organelles contained in the apical tip of their cell. Can be extremely resistant to environmental changes such as temperature or bleach exposure, typically pass through different life history stages in different hosts. Example: Plasmodium (malaria)
Dinoflagellates
Alveolates. two flagella, one equilateral and one longitudinal. Important primary producers . Formed a symbiotic relationship with coral (dinoflagellate performs photosynthesis providing energy for the coral while the coral gives the protist somewhere to grow). Some are parasitic: “red-tides”
Ciliates:
Alveolates. known for their short, numerous flagella called cilia and two types of nuclei. Mostly free living but a few are parasitic. Aid in the digestion of cellulose in cows and termites
Diatoms
Stramenopile. Unicellular, photosynthetic protists that deposit silica (glass like) in their cell walls. Extremely important primary producers in salt and freshwater environments (account for ~20% of the Earth’s total photosynthetic and fix as much biomass as all of the terrestrial rainforests). Went through secondary endosymbiosis in the distant past –> energy production = key to group success.
Coccolithophores
stramenopile which builds a budding exoskelaton of CaCO3 to protect itself from predators. Sequestering of minerals in their shell is highly pH dependent and human ∆s to ocean pH have caused far more CO2 output from these organisms (form a CO2 molecule when they are made).
Brown Algea
Stramenopile. multicellular protist often attaining great size (kelp, seaweed). Important primary producers (food and habitat for other organisms)
oomycetes
Stamenopile. fungi-like absorptive heterotrophs. typically feed off dead organisms through filamentous outgrowths. Some are aggressive plant pathogens causing potato blight and sudden oak death
stamenopiles
Diatoms, Coccolithophores, brown algea, and oomycetes
Excavates
diverse group of single celled, non-photosynthetic organisms, several of which lack mitochondria. This is a derived condition (their ancestors had mitochondria). Euglenids, kinetoplastids, parabasalids, diplomonads, heteroloboseans
Diplomads
Excavate. unicellular protists which lack mitochondria. Include the disease organisms Giardia (one of the most common water born illnesses for humans and other mammals),
Parabasalids
Excavate. unicellular protists which lack mitochondria. Include the disease organisms trichomonas vaginalis.
Heteroloboseans
Excavate. group with two life history stages: one as a flagellated cell and then as an amoeba. Can have members that cause serious infections.
Euglenids
Excavate. distinctive cell surface produced by spiralling strips of proteins, disk-shaped cristae, and flagella with a unique crystalline rod. Common freshwater group. Some are strictly photosynthetic while others are heterotrophs.
Kinetoplastids
Excavate. contain disk-shaped cristae, and flagella with a unique crystalline rod. Typically parasites. Able to change their cell surface to avoid detection by the immune system. Examples: Chagas’ disease, Leishmanaisis, and sleeping sickness
Rhizaria
unicellilar, aquatic, and often have hair-like pseudopodia (a temporary protrusion of the surface of an amoeboid cell for movement and feeding). Can be found in marine or freshwater.
Foraminiferans
Rhizaria. have external porous shells made of CaCO3. Mostly planktonic and very numerous.
Radiolarians
Rhizaria. exclusively marine, many radially symmetrical, and secrete glassy endoskeletons covered in cytoplasm. Many radiolarians harbor other photosynthetic protists such as dinoflagellates.
Plantae
typically photosynthetic organisms, all aquatic with the exception of land plants. Appear to be monophyletic.
Glaucophyte
Plantae. Small group of unicellulat fresh water organisms. They are of special interest becasue they retain some ancestral traits eg. their chloroplasts retain a bit of peptidoglycan between their inner and outer membrane (typical of cyanobact.)
Red Algea
Plantae. marine photosynthetic multicellular organisms red photosynthetic (phycoerythin). Unique in having slightly ameboid male gametes
Chlorophytes
Plantae. morphologically diverse freshwater (a few can be marine or terrestrial) group. Referred to as green algae, posses chlorophyl a and b, store photosynthetic products as starch. Looked at as a proxy for how early plants began.
Unikonts
include fungi, animals, and slime molds. _____________
Opisthokonts
Unikont. Include animals, fungi, and choanoflagellates. All posses a posterior single flagella as opposed to multiple anterior flagella. Choanoflagellates often form colonies and these colonies are thought to be the origin of multicellularity/first animals
Amoebozoans
protists characterized by their thick pseudopods (fake feet) and amoeboid body morphology. Gain nutrients by phagocytosis.
Loboseans
amoebozoans. exclusively single celled organisms that live within a hard covering that is either secreted or glued together from sand grains. Slime molds are surprisingly diverse.