Arthropods Flashcards
What is the most succesfful and diverse group of animals?
Arthropods
85% of all species
Most dominate in fossil record
Where can arthropods live
Cosmopolitan (air, land and water)
The body of arthropods is ______ and possess _______
Segmented
Jointed appendages for feeding and movement
Phylum Arthropoda unique characteristics
- Body epidermis is composed of chitonous exoskeleton which is jointed, segmented and hardened/sclerotized
- lack locomotory cilia in larval and adults stages
Important Arthropod groups include:
- Insecta: anta, dragonflies and bees
- crustaceans: crabs, lobsters and shrimp
- arachnids: spiders and scorpians
- myriapods: centipedes and milipedes
Tagmatization
Body divided into segments with specialized functions
Ex: insecta and crustaceans have 3 distinct body segments (head, thorac and abdomen)
Arthropods possess __________ made up of ______ which is shed periodically
Jointed, flexible exoskeleton
Chiton
Reason for success of the members of the phylum
Exoskeleton is secreted by what:
Epidermal cells
In terms of body plan and development, arthropods are ________ and ______
Bilaterally symmetrical and triploblastic
Body organization of arthropods is of an _____ level
Organ-system
4 layers of exoskeleton
- Epicuticle (outer layer)
- Endocuticle (differ among species)
- Epidermis
- Basment membrane
Exoskeleton is similar to shells of molluscs but differ in
Function and composition
In addition to protection, exoskeletong plays a role in
Movement and composition
Sclerotization/ tanning
Hardening of the cuticle by cross-linking of proteins of the epicuticle
Epicuticle
- Waxy, made up of layers of lipoproteins, underlaid by layers of lipids
- water impermeable (resistent to water loss by dehydration)
- disadvantage: can’t function in respiration directly
- thin, composed of 3% of wrthropod exoskeleon
Procuticle
Forms bulk of arthropod exoskeleton
- exocuticle
- endocuticle
Made largely of chitin (polysaccaride) and proteins
Biotechnological application of chiton
-Chiton has unique characteristics of strength, non-allergenic and biodegradable thus, it is applied as follows:
- dissolved and reformed into fibers for fabrics abd surgical sutures
- making capsulated therapeutic drugs
- proposed for yse as plastic wrap for processed foods: alrhough not digestible by vertebrates
- as food additive: reduce calorie and cholesterol uptake
- purifying agent: high binding potential was harnessed for removing organic and inorganic wastes from table water and sewage purification
The major body cavitt of an arthropod is a
Hemocoel forming the circulatory system
The true coelom of arthorpods is
Greatly reduced to the spaces of the genital and excretory organs as it is encased in a rigid exoskeleton
Arthropods have a complete _____ system
Digestive
With mouth and anus
Excretory organs of arthropods
Green glands, malpighan tubules, cocal glands
Nervous system of arthropods
Similar to annelids with a dorsal brain connected with a nerve ring to double ventral nerve cord
Sensroy organs of arthropods
Eyes (simple or compound), chemo-and tactile receptors, balancing and auditory organs
Molting
- process of shedding of exoskeleton and or cuticle lining the inner gut. Also called Ecdysis
- ensures growth and decelopment among arthropods
- involves secretion of new cuticle underneath the old one by underlying epidermal cells, followed by partial enzymatic degrading of old cuticle which is released.
- newly secreted soft cuticle ensures growth of the arthropod prior to hardening
In crustaceans, secretion of new cuticle requires _____
Cryptocyanin (substance evolved from hemocyanin, copper based protein which function in oxygen transport
Motling is regulated by neural and hormonal controls known as
Ecsysteriod hormine
Motling is induced by release of _____ in head of crustacean or by induction of ________ in the thorax of insects
Y-organ
Prothorax gland
Morling in crustaceans is inhibited by _____ until next molting period is reached
X-organ
Nervous and muscle control in arthropods
Nervous control is theough complex network of axons (slow or fast) which innervates muscle fibers os most parts of the body
Strength of muscle contraction depends on the rate at which nerve impulses are transmitted to the fibers
Arthropods citculstory system
- open circulatory system with the oxygenated blood moving throguh a series of sinuses and finally being drawn back into the heart through ostia as the heart expands.
- dorsal artery carey oxygenated blood theough hemocoelic channels (unshaded vessels) for gaseous exchange between lungs abd tissues blood to the heart
- ventral artery collects deoxygenated blood (shaded vessels) to the gills to be oxygenated and retuned to the heart via the pericardial sinus
Arthropod respiration
Specialized structures
- fills in aquatic forms
- trachea or book lungs in terrestrial forms
Artrhopod ocelli
Simple eyes
- appear like a small cup with light sensitice surface covered by a lens
- it is supported by light absorbing pigments made up of vitamin A derivates suppeoted by a protein
- do not form images
- common in lower invertebrates
Arthropod compound eyes
- Image forming
- May be present alongside the ocelli
- made up of many individual units reffered to as Ommatidia
- gives a broad visual field
- compound eyes break down an image before it reaches the retina, thus, each ommatidia forms a small part of a single image
- common in bees, ants, cockraiches and crabs
Insects may be composed of many thousands of ommatidia which are oriented….
Each oriented in a slightly different direction from each other owing to the convex shape of the eye
5 parts of ommatidia
- Fixed-focus lens or cornea: focuses imsge far and near
- Underlying gelatinous Crytalline cone: serves as lens in some insects and majority if crustaceans
- Cylindrical cells/collars: contain shielding pigment; optically seperates each ommatidium from surrounding ommatidia
- Retinular cells: series of ~8 photoreceptors each containing light-sensitice pigment
- Neural cartridge: situated at the basal end, contains a cluster of neurons which recieves information from the retinular cells and send action potential to the optic ganglia for processing
Sexual reproduction in arthropods
(MOST COMMON)
- Internal fertilization: common in terrestrial species
- most are gonochoristic (seperate sexes)
- hermaphroditic occurs in few species mostly sedentary and parasitic ones
- development is usually indirect through larval stages
- parental care is often well marked in many arthropods
Arthropod asexual reproduction
Parthenogenesis: development of offspring from unfertilized eggs. Common among the class insecta, branchiopoda and some freshwater copepods; some of these members lack males
Mating partners in arthropods locate themselves via
Release of certain chemicals called pheromones
Spermatophores
Are specialized containers used by males of kany arthropods such as, insects, mites, copepods and scorpians, for indirect transfer of sperm to the female
Fertilized eggs of arthropods undergo varying stages of development which could be
Difect or indirect
95% of all arthopods are contained in a superclass…..
Crustacea and two other classes arachnida and insecta
Taxonomic classification of arthropods is mainly on differences in appendages which include:
- number of appendages
- distribution of appendages
- embryonic origin
- form and function of appendages
Subphylum Trilobitomorpha
- called 3 lobed arthopod
- possess two anterior-posterior furrows which divides the body into 3 regions (2 lateral, 1 central)
- no living species
- fossil evidence shows some walked, some burrowed and some swam
Triolibitomorpha body forms
- body generally flattened ventrodorsally and divided into 3 parts: cephalon, thorax and pygidium
- anterior and posterior sections are covered with unjointed sheet of exoskeleton known as carapace; covering the metamerization of these segments
- possess compound eyes made up of many ommatidia
- presence of metameric segments shows a primitive form of arthropods
Subphylum Chelicerata unique characteristics
- most possess claws
- no antennae or manibles
- lack distinct head
- body dividible into 2 portions (cephalothorax or prosoma and opisthosoma or posterior abdomen)
- furst pair of preoral appendages called chelicerae on the prosoma are modified for feeding
- first anteior segment lacks appendages
- second anterior segemnt possess a pair if appendages (chelicerae) adjacent to mouth for grabbing and shredding food
Superclass of subpylum chelicerata
And its two classes
Merostomata
Arachnida: spiders, mites, ticks
Pycnogonida: non segmented body
Superclass Merostomata unique characteristics
- appendages present on opitosoma are flat and adapted for gaseous exchange (book gills)
- terminal part of the opistosoma (posterior segment) possess a long pointed spike known as a telson
Merostomata possess how manu living species?
4, most are extinct
Horeshoe crab (Limulus polyphemus)
Members of the merostomata are all _____ and localized where?
Marine
Seas of eastern north america, indonesia and south east asia
Horeshoe crab use what part to aid in burrowing
Telson
Searching for mucro animals for feeding
Horsehoe crab burrowing helps to
Aerate the soil
Biotechnological application of the Horseshoe crab
- blood components used as bacterial screening tool: components of blood serve as a diagnostic tool for screening injectable pharmaceutical drugs for bacterial endotoxins
- also serves as a diagnostic component for detectinf diseases like meningitis, gonorrhea and others
- vision research: studies on eyes of horseshoe crabs provided insights for us
- important food source: larvae of horse shoe crab released on the sea surface are major food source for kigrateoy shore birds like red knots
Carapace
The head and thorax of merostomata are fused into a single functional unit, the prosoma or cephalothorax and are covered with a single, unjointed sheet of exoskeleton called a carapace
What is the only chelicerate with compound eyes
Hore shoe crab
Generic evidence reveals that the prosoma is similar to what
The hesd of other arthropods
How many appendages do Merostomata have?
6
All 6 pairs are found on the prosoma
First pair modified for feeding
Other 5 pairs are for walking (Pedipalp) and last pair has claws
Distinction of Pedipalp of male and female Merostomata
Males: use it for grasping feamle during mating
Female: same as other appendages
The first 4 pairs of walking appendages of merostomata are modified at base into ____
Toothed food grinding surface known as gnathobase
The fifth pair of appendages in merostomata are modified for
Digging mud burrows and cleaning the gills
Chilaria
Small, hairy appendages present at the last sefment of the prosoma; used for food crushing and transfer of food to the mouth
Opistosoma bears……
Also known as the abdomen bears 6 pairs of appendages; first for reproduction, other kodified as gills
Also bears gill flaps which contain numerous openings (150) book gills; function as surfaces for gas exchnage and channel for blood circulation
(Merostomata)
Class Arachnida
Habitat and examples
All 70000 described speicies are terrestrial; earlier memebrs were marine
Include spiders, ticks, scorpians
50% of arachnida are
9000 species are
Spiders
Mites and ticks
Spider feeding
Carnivorous; prey mainly on insects; maning then a good biocontrol agent
Class Arachnida body
Body divided into 2: Prosoma (fused head and thorax) and opistosoma
Prosoma bears carapace
Arachnida eyes
- spiders possess 0-4 pairs of simple eyes present on prosoma (4 most common)
- lack compound eyes
Arachnida appendages
6 pairs of appendages
- first pair=chelicerae for fedsing
- second pair=pedipalps for locomotion, defense and reproduction
- last 4 pairs for walking
Basal segment of pedipalp in arachnida contain
Maxilla for mechanical separation of food
The prosoms and opistisima of arachnia are connected by a stalk called
Pedicel thst allows for flexibility of the abdomen
Ticks and mites possess fused
Prosoma and opistosoma
Arachnida respiratory organs
- tracheae (small species)
- book lungs or book gills= flattened respirateoy surfaces in the abdomen, they are connected to the outside by means of openings called spiracles
-spiracles close after each breathe to acoud dehydration
Among small sized species of arachnids, spriacles may…
Lead into the tracheae (collection of tubules); thus gaseous exchnage not connected with circulatory system; trachea leads directly into body tissues
Spinnerets
Possessed by spiders, secrete silk proteins which serves several purposes
- scaffold for climbing
- escape danger
- webs for trapping prey
- egg sax that to protect embryo, making homes or mating
Class arachnida sexes and reproduction
Sexes separate (dioecious)
Mostly oviparous and courtship before mating. In some species, the make transfers sperm to the female in a spackage or spermatophore
-usually lay yolky eggs, which hatch into immature young that resemble adults
Arachnid diverse feeding styles
Omnivorous Carnivorous Herbivores Fungivores Parasites