Week 22 Flashcards

1
Q

What is the study of birds called?

A

ORNITHOLOGY.

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2
Q

How many species of birds are there?

A

c. 9,800 species today- changes all the time. Terrestrial vertebrates- which are incredibly diverse.

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3
Q

Smallest bird-

Largest bird-

A

Bee hummingbird- 1.5g

Ostrich- 154kg

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4
Q

What is plumage and what are the aspects of it?

A

In the way they look.
a bird’s feathers collectively.

Extravagant or drab

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5
Q

Where do birds inhabit?

A
Inhabit about every habitat in the world. Some have become adapted to urban life. 
Deserts
Polar ice 
Mountains 
Rainforests
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6
Q

What are the mating systems of birds?

A

Polygyny
Monogmay
Polyandry
Polygynandry

Some individuals pair for life.
Sometimes one male mates with multiple females.
Sometimes females mate with males but this is rare.

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7
Q

What does Bills relate to?

And what are some examples?

A
Bills relates to the type of foraging these species do. 
Probing 
Sifting
Crushing
extracting 
shredding 
hammering
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8
Q

how are birds classed?

A

Phylum: Chordata
Subphylum: Vertebrata
Class:Aves

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9
Q

What does the phylogenetic tree tell us about the birds common ancestor?

A

Shows evolutionary relationships of species which share a common ancestor. It is these last groups of reptiles that the bird’s aves are most closely related to. That is they diverged from a common ancestor more recently than from any other group. So all of these vertebrates represented on the phylogenetic tree have legs. Their common ancestor had legs and this means they are all what are called tetrapods. All but the amphibians also have what are called amniotic eggs. Which makes them amniotes. They all develop through an embryo that is enclosed within a membrane called an amnion. The amnion surrounds the embryo with a watery substance and is probably an adaptation for breeding on land. Amniotes developed directly into a typically terrestrial form, that has limbs and a thick stratified epithelium rather than first entering a feeding laval tadpole stage, which is followed by Metamorphosis, as in the case of the amphibians. You can continue finding features shared by some, but not all of the classes represented here. Feathers is what makes birds unique- they are a modification of the outer skin, so they are this exclusive feature found in all bird species. Archaeopteryx which is the earliest representation of the birds so far discovered that this feather characteristic has been apparent.

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10
Q

What is the Archaeopteryx?

A

•Very early avian
• Late Jurassic Period
• 155–150 mya
Its the imprints of feathers on the fossil remains, extraordinary well preserved. In terms of size and shape it is not that dissimilar to a magpie.

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11
Q

What are the classic reptilian features that the archaeopteryx has?

A
  • -It had a toothed beak
  • -Wing claw
  • -Long tail that contained many vertebrate, so it had bones in its tail.
  • -Also had feathers, this avian characteristic
  • -Was probably a strong running terrestrial bird that could leap into trees, jump among large branches, and perhaps make short flights between trees.
  • -Been agreed that they were capable of gliding and perhaps weak flapping, but not of long sustained flights that we associate with many species today.
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12
Q

Although there is huge diversity in the avian world, all birds have a particular characteristic that defines them- What is this feature?

A

Feathers.

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13
Q

Where did the feathers evolve from?

A

Which evolved from reptilian scales and birds still have scales on the lower parts of their legs and feet. Compared to scales, feathers are filamentous, soft in texture, flexible, lightweight structures. Apart from being a good diagnostic feature, feathers are such extraordinary evolutionary advances that alone need explanation and they contribute so much to the success of birds.

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14
Q

What are the general characteristics of feathers?

A
  • -Made from the protein keratin- beta keratin which is similar to but different from the Alpha keratins that are found in the skin of all vertebrates.
  • -Light, strong & waterproof-
  • -No. feathers roughly constant within a species- although they do tend to have more in winter for those species living in cold environments.
  • -But there is huge variation in the number of feathers found in different species. Ranging from less than 1000 in some of the hummingbirds. Up to maybe 25,000 in these swans- a huge proportion found in the head and neck of the swan as they spend a chunk of their year in the Arctic conditions that necessitate great insulation.
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15
Q

What are the five main types of feathers?

A
  • Contour
  • Down
  • Semiplume
  • Filoplume
  • Bristle
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16
Q

Where are contour feathers found?

A

Cover the whole body except for the beak and the scaled parts of the legs and feet.

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17
Q

What are the features of contour features?

A
  • Give the bird its characteristic smooth, round shape- streamlining the body and therefore increasing the efficiency of flight.
  • Flight feathers (remiges) and tail feathers (rectrices)-
  • First level of defence- on outside of the bird so provide the first level of defence for the bird and protection from physical damage that might arise from vegetation or blowing grit. They also shield the skin from the potentially harmful rays of the sun.
  • Visual colouring- varied and often eye-catching colours of birds are also mostly related to the contour features. Its those that are the coloured feathers that generate the plumage characteristics that we use to identify birds and that they use in their lives to, for example attract mates.
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18
Q

How do the contour feathers get there colours?

A

protein pigments

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19
Q

What are these protein pigments?

A

Melanins (made by bird): black/grey/tan.

Carotenoids (in diet): red/orange/yellow- not generated by the bird itself but is collected in their diets.

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20
Q

What is the intensity of the contour feathers dependent on?

A

The intensity of these colours is therefore dependent on how much carotenoid is being gathered from the food that the bird is catching. As a consequence, the intensity of these colours can be a good and accurate reflection of how good a bird is as a forager. That means that in some species it’s these reds and oranges and yellows that are used by females to assess the quality of a potential male partner.

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21
Q

Why is ultraviolet reflectance important?

A

Ultraviolet reflectance is also important- in the last couple of decades, its also become apparent that the feather colours of birds, especially blues and violets are rich in ultraviolet reflectance. Birds have a fourth UV sensitive cone in their retinas, which allows them to see ultraviolet light. And ultraviolet condition therefore plays a vital role in bird behaviour. Birds seem to look rather drab to us, but with this ultraviolet sensitive cone. Birds see the image on the left. All this additional signalling.

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22
Q

Many of the brightest colours in birds are structural colours that result from the physical alteration of ______ ______

A

incident light.

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23
Q

What are some additional structures of contour structures?

A

Reflectance- blue/green - produced by the reflection of specific wavelengths of light by different sized particles of melanin contained within the contour feathers.
Interference-iridescence - result from the interference of light waves reflected from the outer and inner surfaces of hollow structures within the feathers.
The perception of interference colours depends on the angle at which the bird is viewed. Iridescent colours therefore kind of flash on and off as the birds changes its position.

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24
Q

What is the structure of a contour feature?

A

Rachis
Barbs
Barbules and hooks

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25
Q

Rachis-
Barbs-
Barbules and hooks-

A

Rachis: hollow, central shaft providing backbone.

Barbs: extend either side; site of most pigments.- have small barbules radiating out from them. Are the site of most of the pigments.

Barbules & hooks: create stiffness & flexibility; fasten feathers to one another. These little barbules and hooks link up together to create both a stiffness and a flexibility in the outer surface of the bird. Effectively they’re fastening their feathers to one another to create this continuous, streamlined surface. Preaning is partially used to restore any interlocking of these barbules and hooks that comes undone.

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26
Q

Type II: down feathers

A
  • Smaller than contour feathers-
  • Lack barbules and hooklets: not zipped together meaning that they are not locked together.
  • Soft and fluffy
  • Insulation- they are found closest to the skin of the bird, providing this insulating layer of warmth.
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27
Q

Type III: semiplume feathers

A
  • Intermediate in shape between contour and down
  • Found between contour feathers
  • Help with insulation
  • Help keep form- fill out the body contours, it helps to keep the form in general.
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28
Q

Type IV: filoplume

A
  • Hairlike: few barbs at tip
  • Covered by other feathers- not exposed, never found alone and are found next to contour feathers.
  • Pressure and vibration receptors- each flight feathers for example might have 8 to 12 filoplumes associated with it. also , unlike other feathers, they have no muscles at their base. But their follicles are endowed with abundant nerve ending, which are sensitive to changes in pressure and vibration. Effectively they are operating as these pressure and vibration receptors.
  • Help keep other feathers in order- act as sensory organ for keeping the other feathers, particularly the contour feathers in place and adjusting them in flight and for insulation and during things like bathing.
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29
Q

Type V: bristle

A

Whereas all birds have the other four types only some have bristle feathers. They are stiff feathers with a few barbs near the base. In those birds that have them, they tend to be found around the eyes and the mouth. They function as sort of guard hairs, protecting those vital organs. They might also act as some sort of sensory organ a bit like the whiskers of a cat. And in some birds, particularly those that catch insects on the wing, they might act as a funnel to funnel those invertebrates towards the beak, enhancing foraging behaviour.

  • Stiff, with a few barbs
  • Occur around eyes and mouth of some birds
  • Protective/filters
  • Funnels for insectivores
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30
Q

What are sociable weavers?

A

a species found in southern africa, each pair of sociable weavers, sometimes with a few little helpers, builds its own nest chamber. But these sociable weavers build collectively together and form these enormous colonies. They are so big sometimes, so many pairs contributing to this overall structure, each with their own next cavity that the weight of the nest can bring down trees or telegraph poles. Because there are lots of nests in the same place, this makes them really attractive to predators like these snakes who come searching for eggs or young chicks. Now the sociable weavers can attempt to chase off snakes, but they don’t have anything other than the ability to mob and yell at them to try and get rid of them. They are aided in getting rid of snakes by pygmy falcons. Often you find a pair of pygmy falcons nesting in one of the chambers in these huge sociable weaver colonies and they eat snakes so they act as a protective force to the sociable weavers. Interspecific interaction.

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31
Q

what are the functions of feathers?

A

provide colouration

Crypsis
Attraction
Insulation
Flight

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32
Q

What is Function I: crypsis?

A
  • Blending into background to avoid detection
  • Predators sneaking up on prey
  • Prey escaping predators
  • Behavioural traits to match

Cryptic colouration helps hide or camouflage the bird, lowering the likelihood that the individual will be noticed. Its helping them blend into the background to avoid detection for some reason. Some predatory birds have cryptic colouring that guarantees a more dependable approach to prey. For example the typical white colouring of the head, neck and underparts of many fish eating birds helps them to remain undetected by fish looking up into the bright sky. More commonly cryptic colouring is used to escape the attention of predators to avoid being eaten by something nasty. For these protective colourations to be effective, it is sometimes important for certain postures to be adopted and for the bird to remain immoble.

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33
Q

What is Function II: attraction?

A
  • Attract mates
  • Males usually brighter than females
  • Sexually selected trait
  • Brightest males picked by females

Some birds have plumages designed to stand out from background and show off. They are generally used to attract mates. And in many species, its the males who are brightly coloured. That is because often it is females who are the choosy sex and males who are trying to attract selection. As such, this is a sexually selected trait found in one sex and not the other. So many species that have brightly coloured males, females are actually rather drab brown. Theres no need for them to show off. And they are actually better off being less conspicuous and less likely to be spotted by predators. In these species where males are displaying their wares through their plumage, its often the brightest coloured males who are picked by females. That may be inherent quality of the male, or it may be a good forager and able to collect more of these carotenoids that form the reds and oranges and yellows in the plumage.

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34
Q

What is Function III: insulation?

A
  • Primary force driving evolution of feathers
  • Vital for endothermy (body temp. c. 40°C)
  • Denser plumage in winter
  • Fluff out feathers in cold conditions

Insulation is believed to be the primary force driving the evolution of feathers. As endothermic organisms birds maintain a constant body temperature, which can be as high as 40 degrees. The insulation provided by the feathers greatly reduces the amount of energy required to keep up the body heat. And in some birds they moult and have a denser plumage in winter, particularly those living in very cold environments. And just like with crypsis for example then these feather characteristics can be enhanced by behavioural traits. For instance, if you look out on a cold morning in winter here in the UK, you might see a bird looking rather fluffed up as it deals with those early morning temperatures.

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35
Q

what is Function IV: flight?

A

Feathers are also important in flight, but initially evolved for insulation. So flight followed that initial evolution of veined feathers. Early forays by gliding and weak flapping flight probably fostered additional changes in feather form and function on the wings and the tail particularly. Feather design nowadays gives birds the strength, flexibility and rigidity to support themselves in the air, which of course provides very little support itself. Theres all sorts of characteristics about flight, not just feather form and function.

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36
Q

What is the most basic feather care?

A

Preening

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37
Q

Why is preening important?

A

•Restore feather structure, clean plumage, rearrange feathers- depending on the type of maintenance of the bill is used in one of two ways. Either the feather is worked between the mandibles of the bill with a sort of nibbling action. Which is probably important for restoring the relationship between the barbules and the hooks. Relocking those feathers contour feathers together again and for removing dust particles and stale oils. Second way of manipulating in preening is where bills are wiped across the surface of the feather and thats probably important in rearranging them a little bit in drying them out or particularly in the application of preening oils.

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38
Q

Why is applying oils useful for feather care?

A

•Apply oils from the uropygial gland for waterproofing- these oils come from something known as the uropygial gland and they are crucial for waterproofing the feathers. Not all parts of the body can be reached by a bird’s beak, especially if that beak is a particular shape. Sometimes the head and neck are rubbed with the feet instead of the bill.

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39
Q

Why are feathers fundamental?

A

Feathers are fundamental to survival and so feather care is of the utmost importance. Because feathers are inert. And do not have an internal system of nourishment and maintenance. Left untended they would become brittle with age and exposure were it not for regular care, which includes the application of waxy secretions produced by the preen gland.

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40
Q

What is allopreening?

A

In certain species you also see whats known as Allopreening, thats where one individual preens another, either in turn or simultaneously. Studies have shown that although one function of Allopreening is hygienic, reaching bits of the body that an individual cant reach itself. This allopreening may well serve social functions as well.

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41
Q

What is another feather care behaviour?

A

bathing

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42
Q

Why is bathing useful?

A

Bathing is another behaviour that is used for feather care, although it is most commonly carried out with water, some birds such as pheasants, use dust instead

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43
Q

What happens after bathing?

A

After bathing in water, the bird is in a particularly vulnerable condition as its flight efficiency is somewhat impaired. Therefore, it needs a nearby safe place in which to preen itself and to let the feathers dry.

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44
Q

Aside from its role in feather care, bathing may provide a

A

mechanism to assist in body cooling in hot conditions.

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45
Q

What is sunning?

A

In some species you also see this so called sunning behaviour, where the bird settles onto the ground in some relatively protected location. And exposes itself itself to the full sunlight, spreading out its wings and its tail. And often the bird is also fluffing out its feathers in this position. Sunning probably has several functions associated with feather care. One of those is that the sun’s heat is invaluable in restoring and maintaining the shape of the major flight feathers. Also, the interaction of sunlight with the preening oils may be important in synthesising vitamin D. the warmth of the sun may help the spread of preening oil on the feathers or of softening old oil to help their removal.

46
Q

What is anting?

A

And finally, some may stimulate the activity of ectoparasites. Which can then be more easily spotted and picked off by the bird. In some cases theres also this rather intriguing behaviour known as anting. Which involves the bird squatting on the ground in an area where ants are common. The ants then run freely over the bird throughout the plumage, and its suggested that birds exploit the formic acid produced by many ants, which helps remove stale preening oil from the feathers. The ants may of course also be catching ectoparasites and helping hygiene in that regard.

47
Q

Why is moulting useful?

A

Because feathers are subject to damage and wear. Despite all of the feathers care that goes on. Plumage effectiveness is only going to be maintained across time if damaged and worn out feathers can be replaced. And this occurs on a regular basis through a process known as moulting. Moulting is this replacing of old feathers with new ones. Now the number of times that happens per year varies from species to species or even within species, but it occurs at least once a year in all birds. It is an energetically costly process, and of course when you are missing some feathers then flight efficiency or otherwise is pampered. So moulting does not coincide with energetically other costly activities, or those requiring the feathers for good reason. So not when, say the birds are breeding or migrating. It would also be disastrous, of course, if all the feathers dropped off at once and had to be re grown. So if feathers are re places in an orderly predictable sequence and thats particularly true of those involved in flight, so the wing and the tail feathers. Moulting not only allows a bird to replace old feathers, but of course it allows a dramatic change in appearance. In some species where males are brightly coloured to attract females in the breeding season, what you find is that once they no longer need to be attracting females, males mouly into drab, cryptic plumage during the non breeding season and then moult back into their bright colourations at the start of their next breeding season.

Replace old feathers with new ones
1+ times per year
Lot of energy: not when breeding/migrating

48
Q

Feathers replaced in

A

orderly, predictable sequence – especially wing and tail feathers

49
Q

All birds possess …

A

Forelimbs modified as wings. And indeed, they have a particular shape known as an airfoil, which is critical for lift. All birds have feathered tails, that is they no longer unlike their reptilian ancestors, have bones in the tail, they just have feathers. They have a toothless horny beak, again unlike Archeopteryx for instance, they have no teeth because those are very heavy features, so they have lost those in terms of lightening for flight.

50
Q

What is endothermy?

A

Maintenance of constant body temperature. Birds are of course endothermic, so they maintain a constant body temperature.

51
Q

TO be endothermic, what do birds need?

A
  • High metabolic rate
  • Good insulation
  • characteristics shared with mammals
52
Q

What are the advantages and disadvantages of endothermy?

A

Advantage: live in extreme environments- very hot ones or cold ones, which is impossible for many ectotherms.
Disadvantage: energetically expensive- you need to be eating all the time. The metabolic rate of birds is about 10 times that of amphibians and reptiles that need a constant source of fuel, so they have to eat much more.

53
Q

What are some aspects of ancestor bird flight?

A

Ancestor could fly
But, NOT a defining avian characteristic
Some birds have subsequently ‘lost’ flight

The ancestor could fly to some extent, probably only weak flapping flight at best. But this is clearly not a defining avian characteristic. That is because other taxa can fly, think bats for e.g. and also that some birds have lost the ability to fly, like the brown kiwi. So although we think of flight and birds as being synonymous in many senses, its not a defining characteristic.

54
Q
Summarise bird diverisity:
•Diversity
•Taxonomy
•Feathers
•Other avian features
A
•Diversity
–Size, plumage, habitat, mating systems, bills
•Taxonomy
–Reptiles, Archaeopteryx
•Feathers
–Characteristics, types, colouration, functions, care
•Other avian features
–NOT flight
55
Q

Why has gliding evolved and what animals is it seen in?

A

Gliding has evolved on many occasions usually to aid canopy animals in getting from tree to tree. Gliding has particularly evolved among rainforest animals, especially in Asia where the trees are very tall and quite widely spaced. We see gliding in some Chinese gliding frogs for instance, in paradise tree snakes and in gliding ants. Also in flying squirrels, which glide between trees.

56
Q

How do flying fish show gliding?

A

Have also been observed to glide often for up to hundred of metres on the drafts, on the edges of waves, with only their initial leap from the water to provide height. So gliding is pretty common in the natural world.

57
Q

Where has active flapping flight evolved?

A

By contrast active flapping flight has only evolved in four main groups, the now extinct pterosaurs, bats, birds and insects. In many senses birds are the most specialised of these for flight.

  • -Bats
  • -Birds
  • -Insects
  • -Pterosaurs
58
Q

What adaptations do birds have for flight?

A

Birds have a large number of adaptations for flapping flight.

1-feathers
2.lightening

59
Q

What are some features of feathers that help flight?

A

Light
Strong
Streamlined
Flexible

60
Q

Why do the feathers need to be light?

A

much lighter than the scales in which they evolved from.

61
Q

Why do the feathers need to be strong?

A

in order to avoid snapping from all the forces being applied during flapping flight.

62
Q

Why do the feathers need to be streamlined?

A

contour feathers can be locked together to form this ideal aerodynamic streamlined surface for air flow. In producing this smooth uninterrupted outer surface over which air can pass freely. Birds reduce whats known as the drag that would hold them back.

63
Q

Why do the feathers need to be flexible?

A

they give under pressure, thus letting air pass more easily over the body without disrupting the flow and causing excess drag. They are designed to maximise aerodynamic properties in changing conditions.

64
Q

Where does lightening for flight occur?

A

Skeleton
Beak
Reproduction

65
Q

What makes the skeleton light?

A

–Reduced number of bones- unnecessary bones have been lost- no bones in the tail and are now just made up of feathers.
–Bones hollow and lack bone marrow- the remaining skeleton is made lighter than in other taxa because many bones are hollow, they are honeycombed with airspaces and strengthened by crisscrossing struts. They also lack bone marrow which is a heavy substance. Number of hollow bones varies from bird species to species. Diving birds for example dont have any hollow bones bc they need the weight, whie large gliding and soaring birds tend to have many hollow bones. The more efficient flyers seem to have more bones that are hollow.
–Skull lost heavy jaw- birds skull is also extremely light relative to other taxonomic groups. Thats partly due to the elimination of a heavy jaw structure including big jaw muscles and also to the loss of teeth.

66
Q

What makes the beak lighter?

A

–Toothless- so they have a toothless beak now compared to archaeopteryx. Instead of teeth birds have a structure called the gizzard. Its a functional analogue of mammalian molars. A large strong muscular structure used primarily for grinding and digesting tough food, but its much lighter than teeth.

67
Q

What makes the reproductive system lighter?

A

–Organs enlarged seasonally- only develop reproductive organs in the breeding season. Females have just one ovary and it increases by up to 1500 times in size when required.
–Oviparous- that means they lay their eggs as soon as they are fully formed, the vast majority of embryonic development is done outside the female’s body, thus saving her from carrying around the extra weight of developing young inside her.

68
Q

What is a consequence of loosing certain bones?

A

As a consequence of the loss of certain bones, the hollowing of at least some of those that remain and the reduction in the weight of the skull, a pelican for e.g. which is approximately 5 feet long only weighs about 9kg of which the skeleton is only 650g. And we can contrast that to the otter which weighs much more.

69
Q

Although a present day bird has _____ bones than its ancestors. Its skeleton is strong enough for flight.

A

fewer

70
Q

What makes the skeleton stronger?

A

–Fusion of bones to form rigid girders and platforms.
–Wishbone- which is formed by fusion of the collarbones at their base, offers strong structural support for the wings and provides a brace during the powered stroke of flight.
–Keel- And in flying birds the breast bone is fused to a deep keel. This longitudinal ridge of bone. That keel provides an anchor for the powerful flight muscles. The deeper the keel the more powerful the flight of that bird.

71
Q

What muscles aid in bird flight?

A

•Muscles- two main flight muscles attached to each wing.
–Pectoralis (up to 35% total weight)- powers the downstroke of flight, thats the one that generates most of the propulsion. Its therefore portionally huge compared to other taxa.
–Supracoracoideus- its much smaller than the pectoralis muscle and has a tendon which curves around to attach to the top of the humerus. Can provide power to the upstroke if required. But more usually its just a recovery stroke to get the wing back into the upward position for the powerful downward stroke.
–48 other wing muscles- which are more subtle alterations in wing shape and form during flight and especially during takeoff and landing.

72
Q

With a highly developed NS, what are birds main sense?

A

Main sense is sight- rather than smell which we see in most reptiles and mammals.

73
Q

What do birds have to enhance their sight?

A

Large eyes
Large brains-
Ample processing centres- the brain expansion in birds relative to reptiles is of centres related to sight. The processing centres for coordinating visual information received from the eyes. And to be able to process that quickly while they are flying rapidly. That’s in contrast to brain evolution in say rodents, where there’s been more development of the cerebral cortex.

74
Q

What are the wings?

A

highly modified forelimb
Elaborate joints
Fused hand bones
Powerful tendons and compact tiny muscles

75
Q

How do the wings function?

A

The humerus, ulna and radius bones are homologous to those in the limbs of other vertebrates. But they have more elaborate joints which permit the wing readily to change position and angle during takeoff, flight and landing. When outstretched, these joints are strong enough to withstand the wrenching forces created during flight. And that’s helped by having fused hand bones, there are only three digits in the birdwing as opposed to the five found in most tetrapods. That fusion provides strength and rigidity in the outer wing skeleton. And then for their support and manoeuvrability for the large primary flight feathers. Within the wing itself are powerful tendons and compact packages of tiny muscles that can control really in a detailed way the wing positions.

76
Q

Why is the airfoil shape to the wings beneficial?

A

Airfoil in cross section- that is it has this rounded leading edge and a sharp trailing edge. An asymmetric profile is fundamental to generating lift crucial for flight.

77
Q

What is bird flight governed by?

A

Bird flight is governed by the laws of physics, just like aeroplane flight. And that means that there are two sets of competing forces at work .

78
Q

What are the two competing sets of forces?

A

Weight vs lift
Drag vs propulsion

to counteract the downward force of gravity, which is expressed as a birds weight . an upward air pressure force known as lift must be generated. To counter the slowing influences of the forces of turbulence and friction. Which are collectively known as drag. The forward force called propulsion or thrust must be created. When these four forces are in dynamic balance. A bird maintains a level flight at a constant speed. Unlike aeroplanes which generate lift with wings and thrust with engines. Birds use their wings to do both.

79
Q

How does lightening affect weight?

A

birds have of course evolved to be as light as possible. That’s the reason for those lightning adaptations that we saw. Things like having a reduced number of bones, having hollow bones, no teeth, no bones in the tail. And having those reproductive adaptations that all reduce weight and thus the pull of gravity by minimising weight birds minimise the amount of lift that they have to generate.

result of gravity
reduced as much as possible

80
Q

How is lift generated?

A

Now lift is generated by the flow of air over the wings, to understand how this works we have to understand that there is a relationship between the speed of air flow and the pressure created. This is something known as Bernoulli’s principle.

speed of air flow flow relates to pressure.

Further to travel, air speeds up, pressure reduced Wing sucked upwards.
Shorter distance, air slower, pressure increased, Wing pushed upwards.
More curved the airfoil, greater the lift.

81
Q

How does the airfoil shape affect lift?

A

Remember that the wing has this airfoil shape, this rounded front edge, this trailing tapering edge and its this airfoil shape that is so important for the generation of lift. Across that top curved surface there is farther for the air to travel. It therefore speeds up and sir that is travelling faster, generates less pressure.this means that the wing is sucked upwards. Simultaneously air passing underneath this airfoil shape has a shorter distance to travel. The air therefore travels slower which creates an increased pressure. That pushes the wing upwards. So we are getting lift because of this asymmetric shape of the wing with different distances for the air to travel across each of the two surfaces. The amount of lift generated increases with airspeed and with the volume of air deflected over the wing. Moreover the orientation or angle of attack of a wing in a current of air also affects the generation of lift. Increasing the angle of attack, basically making the airfoil more curved increases the amount of lift generated . however if that angle of attack becomes too great, stalling occurs. Birds actually do this deliberately when they are coming into land, because then its beneficial to bleed off that lift and consequently their speed too.

82
Q

What are the two types of drag?

A

Friction

Induced

83
Q

What is friction drag?

A

Consequences of air flow over body
Resistance to anything passing through air.

Friction drag is the drag produced by the flow of air over a body. And this is influenced by uneven surfaces or anything jutting out. Basically from the body so theres resistance to anything passing through the air. That resistance is greater the more projections there are. That means that reducing drag is aided by streamlining.ome of the advantages of the contour feathers is that they can be locked together to produce this streamlined surface.

84
Q

What is induced drag?

A

the second type of drag is induced drag, this is the consequence of lift so we have air passing at different speeds over the top and the lower surface of the airfoil shaped wing. When those two air flows then meet at the wing tip , what is generated is the swirling wingtip vortices. These swirling vortices generate this induced drag, pulling the bird back, holding it back to some extent. These forces, this induced drag is also dependent on the bird speed. So at increased speed there is increased induced drag at the wing tips. And this induced drag produces a downwash, effectively it pulls the bird back a little bit.

85
Q

How can you take advantage of induced drag?

A

But you can take advantage of the induced drag, the wingtip vortices of others if you fly in the right formation. So this is something we see in terms of long distance migrating birds like geese or cranes where we see them flying in this V formation. And thats bc if you fly just behind and just below the bird in front in a certain position, you can take advantage of those swirling vortices of the bird in front. And that reduces your own energetic cost of flying. Its analogous to the slipstreaming you can do in the cycling race if you get in just behind the bike in the front.

86
Q

What is the final force and what does it involve?

A

Propulsion/thrust

  • created when wings flapped
  • majority of force in powered downstroke
  • simple recovery upstroke.

Final force to consider is this forward momentum, propulsion or thrust. This is created by the flapping of birds wings . the majority of the force in most birds is in the powered downstroke that is generated by the pectoralis muscle. The huge muscle in many bird species. In most species there is a simple recovery upstroke that just gets the wing back into the position for that power downstroke.

87
Q

What is the most simplified type of flying?

A

most simplified types of flight is gliding and this illustrates how forces of weight, lift and drag can work. Gliding birds are using their weight to overcome air resistance to forward motion. That means that the heavier the bird is the better they tend to be at gliding, so you really only see gliding or certainly gliding for long distances in large heavy birds. Effectively youre always going to be losing height because as you go along without some other reason to generate that. But the amount of height you are losing while gliding also varies in terms of the distance traveled, so how far you can get for the same height lost basically is dependent on the weight of the bird.

weight used to overcome air resistance to forward motion
Generally used by large birds
Height lost/ distance travelled.

88
Q

What is an example of gliding?

A

So on the left here we have a Fulmar, a seabird weighing about 800g. And in normal conditions it can travel about 8 meters for every 1 metre of height lost while gliding. By contrast the much heavier bald eagle can get about 20 metres for every metre of height lost while it is gliding.

89
Q

How can height be gained?

A

ne way in which height can be gained without going through flapping flight is through soaring.This is maintaining or indeed increasing altitude without flapping. What you need are updrafts of air, which you can then take advantage of and these updrafts can aride in several different scenarios. For instance, there are things called slope soaring. So this is the situation where wind is deflected upwards by the side of objects. Hillside or a mountain side, or even a building. This is the updraft thats often used by human gliders. If you see them along the coastline, making use of these updrafts along the edges of cliffs. Another type of soaring is whats known as dynamic soaring. This is making use of the little localised updrafts when wind meets waves out in the open oceans. So thats just above the surface of the ocean. There are these little updrafts from each individual wave. If you can keep catching those then you can soar for hours at a time with no flapping flight. This is something that the albatrosses are highly adapted to do hours of flying just above the ocean surface with no active flapping flight needed.

90
Q

What is soaring?

A

Maintain or increase altitude without flapping

Slope soaring- wind deflected upwards by side objects

Dynamic soaring- local updrafts when wing meets waves.

91
Q

What is thermal soaring?

A

Thermal soaring: uneven heating of air near ground creates convective currents that cause air to circulate

In terms of soaring theres whats known as thermal soaring, this is because there is uneven heating of air near the ground, which creates convection currents, causing air to circulate upwards. So you end up with these columns of circulating warm air. And thats why birds using these thermals tend to fly in circles. They are trying to stay in this column of upward moving air. This is particularly generated in areas of hot climates. And as thermals are generally not developed over large water birds tend to collect in large numbers in places like Gibraltar where they try to obtain as much height as they can on the basis of thermals to then help them negotiate the straits into Africa with minimal active flapping flight, so they are trying to save energy by getting as high as possible in the thermals before crossing the seas.

Birds in thermals typically fly in circles

92
Q

Why is flapping flight useful?

A

•Adds thrust
•Asymmetric wing actions
•Steering, turning and twisting controlled by numerous muscles.
Flapping flight of course adds thrust this forward momentum to the controlling forces. Each primary feather acts as an airfoil, as does the wing as a whole. And you’ve got these 50 or so different muscles controlling the wing movements: some fold, others unfold it, some pulling up, some pull it down, and still others adjust its exact orientation, allowing precise steering, turning and twisting during active flight.

93
Q

What is hovering?

A

Maintaining stationary position in the air.

Finally there is hovering, birds can achieve this in two ways. Hovering in general means maintaining a stationary position in the air relative to the ground below you. In some large bird species this is done by wind hovering.

94
Q

What are the types of hovering?

A

Windhovering- fly into headwind and use thrust to fly slowly.

True hovering-wing airfoil shape in up and down stroke

95
Q

What is an example of hovering?

A

So in this Osprey for example what it does is it flies into a headwind that exactly the pace needed to maintain a position above the surface below. And its doing that because in the case of the Osprey, its looking for fish to feed on. So its trying to hold a stationary position in the air to be able to have a clearview and lookout for its lunch. The real experts, those species that can manage true hovering, so even in still air. well , the largest of those in the world is something called a pied kingfisher in Africa. But the real experts are the hummingbirds. And they achieve this because, unusually they have power in both their downstroke and their upstroke. They can flip their wing over such that theres an airfoil shape both times round. And they can generate perhaps 52 beats per second, which means that not only are they flapping very very fast, but both the upstroke and downstroke generates power.

96
Q

What are the elements of wing shape?

A

Wing shapes- vary substantially among species. And the shape of the wing is an important factor in determining the types of flight of which the bird is capable or best adapted. Different shapes correspond to different tradeoffs between beneficial characteristics, such as speed, low energy use, gliding ability, and aerial agility. If we look at the planform of the wing, thats the shape of the wing as seen from below. It can be described in terms of two parameters.

97
Q

What is the first of the two parameters?

A

First of the two parameters is known as wing loading- thats the ratio of weight to wing area and it determines the costs of flight. Birds with a low wing loading have a large wing area for a particular body weight. The other extreme we have birds with a high wing loading that is a relatively small wing area for a particular body weight. Birds with low wing loading tend to need less power to sustain flight. And these species tend to fly more slowly and that are more maneuverable than those with higher wing loadings.

Low wing loading - Large wing area for a particular body weight.

High wing loading - Small wing area for a particular body weight.

98
Q

How do you calculate wing loading?

A

Body weight/ wing area

99
Q

What is an example of wing loading?

A

Higher wing loading can also be a problem in terms of taking off. You’ve got to get this big weight off the ground with a relatively small wing area. For example swans which are taking off from ground level- the only way that they can do this is to run along the surface of the water while flapping, they are trying to generate sufficient speed to get enough lift to get off the ground. Vultures are also species with high wing loadings but they stand on the top of things like cliffs and just launch themselves into space and that downward momentum and that increase in speed as they plummet results in lift sooner rather than later.

100
Q

What is the second parameter?

A

The second parameter, in terms of wing shape, is called the aspect ratio. Thats the ratio of the wingspan to the wing area. Generally the higher the aspect ratio that is longer narrower wings , the lower the induced drag produced by the wing at a given speed, remember induced drag is generated at the wing tips due to the different speeds of air passing over the top and bottom surfaces of the airfoil shaped wing. Most bird wings can be grouped into four types of aspect ratio. Of course it’s more like a continuum but its easier to categorise in general senses. Particularly when we are interested in thinking about what particular aspect ratios generate in terms of characteristics for flight.

101
Q

How do you calculate aspect ratio?

A

Wing breadth squared/ wing area

102
Q

What is the second type of wing?

A

high lift wings

103
Q

What are high lift wings?

A

These have a moderate aspect ratio. They’re also pretty broad wings, just like the elliptical ones, but somewhat longer. They are also heavily cambered and these wings are particularly good for carrying a heavy load, so a big body mass, whilst minimising the energy needed to fly. These high lift wings are particularly common in terrestrial soaring species. Things like the Eagles, the vultures and the storks. These are the species that are particularly making use of thermal soaring. Remember those columns of circulating warm air rising from the surfaces? And these high lift wings are excellent to allow broad turns, as the birds circle inside these thermals and to catch those thermals and lift the birds to higher and higher altitudes. These high lift wings also suffer from lots of induced drag, so we often see wing slots in these species too.

  • Moderate aspect ratio
  • Heavily cambered
  • Carry heavy load, but minimise energy
  • Terrestrial soaring species (e.g. eagles, vultures, storks)
104
Q

What is the third type of wing?

A
  • Relatively high aspect ratio
  • Almost flat profile
  • Aerial feeding (e.g. swallows, swifts)
  • Long migrations (e.g. shorebirds)

A third type of wing is the high speed wing. This is rather different, these have a relatively high aspect ratio. They have an almost flat profile, but you’ve got this pointed slender wingtip. So theres no need here for any wing slotting because already the level of induced drag in minimised. And they have a swept back look too. These high speed wings are particularly suited to species that feed aerially chasing invertebrate prey up in the sky. So we see high speed wings in, for example, swallows and swifts. These narrow tapering wings can also be flapped rapidly to generate lots of speed with minimal drag. That means they are also good for long migrations, so we see this type of wing shape in many shore birds, which migrate big distances between breeding and wintering grounds.

105
Q

What is the final wing shape?

A
  • High aspect ratio
  • Narrow, pointed shape reduces drag while gliding
  • Sea birds in windswept environments (e.g. albatrosses).

The final wing shape is this soaring wing shape. They have a very high aspect ratio, that these long thin tapering wings again because of that narrow wingtip, theres no need for wing slotting. And these are particularly good at reducing all sorts of drag whilst gliding. We therefore see these soaring types of wing shape, particularly in seabirds in windswept environments. Classic example being the albatrosses- which are experts at dynamic soaring. Picking up these little updrafts from individual waves out in the open ocean, constantly adjusting and making use of those soaring options close to the ocean surface. And these soaring wing shapes are ideal for that way of life.

106
Q

What is flight often used for?

A

Flight is often important for feeding, take these hummingbirds, for e.g. they have to move from flower to flower, be able to extract the nectar constantly, and move on. Others are feeding on invertebrates that they are chasing in the sky. Still others are using flight to be able to spot prey down on the ground before diving down to pounce upon it.

107
Q

What is flight also used for?

A

Flight is also sometimes important in courtship. Here we have a male and a female in perfect synchrony during a courtship flight. While they are choosing a partner. And flight opens up travelling for huge distances. There is no way that a mammal of an equivalent size could travel such massive distances as exhibited by this Willow warbler. They weigh only a few grams and yet annually they make a migration from Alaska to Sahara, 10,000 kilometres or so. That is only made possible through flight.

108
Q

What are aspects to under water ‘flight’?

A

Increased weight
Reduced buoyancy
Smaller wings

There are some species which spend much more time under water like these penguins. In these species there is increased weight compared to those birds that fly in the air, you don’t need to overcome gravity compared to those flying in air, and indeed you might want reduced buoyancy so you don’t have this same hollowing out of the bones, for instance. These species flying underwater also have much smaller wings. They don’t have to generate lift and thrust in air. Instead they are much more useful for steering underwater.

109
Q

Why have some species experienced loss of flight?

A

Expensive
No need if reduced predatory risk

Not all modern day birds fly, there has secondarily been the loss of flight in some species, like the brown kiwi. The reason for that is that flight is incredibly energetically expensive. Thus if you can get away without needing it, then you can save that energy for other parts of your life. For instance, if you live on an island where there are no terrestrial predators. Then theres no need for flight to escape that threat. In which case there is the possibility to have evolved the loss of flight and saving of all that energetic output.

110
Q

Summarise flight:

  • Adaptations
  • Mechanics
  • Types of flight
  • Wing shapes
  • Functions
A
•Adaptations
–Feathers, lightening, strengthening, brain and sensory organs, wings
•Mechanics
–Weight, lift, drag, thrust
•Types of flight
–Gliding, soaring, flapping, hovering
•Wing shapes
–Elliptical, high speed, soaring, high lift
–Feeding, courtship, migration, diving
111
Q

What is the first wing type and why is it beneficial?

A

Elliptical.

First wing shape we can think about
•Low aspect ratio- their broad chunky wings if you like
•Good manoeuvrability (e.g. woodpeckers, forest raptors)- birds with these types of wings have good manoeuvrability, they can flap them very relatively fast. They can manoeuvre well. That means they are extremely useful in tight environments, so when theres lots of vegetation around, in say woodlands or forests. So birds like woodpeckers and forest raptors have these elliptical wing shapes.
•Fast take-off (e.g. pheasants, grouse)- this type of wing shape also allows fast takeoff from the ground. Thats especially useful for species that forage predominantly on the ground, but which are vulnerable to terrestrial predators. Think here things like pheasants and grouse, as they move along the ground looking for food. If they spot a fox, they need to get out of the way off the ground quickly, and these elliptical wings are best for that. The trouble with these broad wings with a low aspect ratio is that they generate large amounts of induced drag.
•Slotting- Birds with elliptical wing shapes have therefore evolved this, slotting on the wing tips. This is the gaps that you see in the diagram between the feathers. That allows air to pass through and dissipate and the bending of the feathers leads to the displacement of the wingtip vortices. Spreading it over a greater area of the wing and leading to reduced induced drag.