Sensation & Perception Final Review Flashcards
What form of energy does each sense capture?
Photons, air pressure, chemicals, etc.
Vision: Electromagnetic
Audition (Hearing): Mechanical forces
Vestibular (Sense of balance): Mechanical forces
Proprioception (Sense of limb positions):
Mechanical forces
Touch
How can the different forms of energy be described?
Dimensions? Units? Etc.
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The quantitative relationship between physical energy and sensation:
psychophysics
How does physical energy in the environment become transduced into neural signals within the brain?
sensory physiology
How are patterns of sensory stimulation interpreted as meaningful events?
psychophysiology
psychophysics
The quantitative relationship between physical energy and sensation
sensory physiology
How does physical energy in the environment become transduced into neural signals within the brain?
psychophysiology
How are patterns of sensory stimulation interpreted as meaningful events?
Dualism:
The idea that the mind has an existence separate from the material world.
Mental phenomena are non-physical.
Rene Descartes (1596-1650):
- Defender of dualism
- Father of modern philosophy -Invented Cartesian coordinate system (X, Y, & Z)
- “I think therefore, I am”
Rene Descartes
(1596-1650)
- Defender of dualism
- Father of modern philosophy -Invented Cartesian coordinate system (X, Y, & Z)
- “I think therefore, I am”
Materialism
The idea that only matter and energy exist.
The mind is not separate from the body.
Most psychologists are modern materialists.
Gustav Fechner
1801–1887
Invented “psychophysics” and is often considered to be the true founder of experimental psychology.
Struggled with the mind/body problem
Worked to exhastion. Went blind looking at the sun. Depression. Spent 3 years alone. Experience a “miracle” when vision returned. Struggled with mind body Also had some crazy ideas.
Believed in panpsychism: The idea that all matter has consciousness.
Wrote Nanna, or Concerning the Mental life of plants
Ernst Weber
(1795–1878) discovered that the smallest change in a stimulus, such as the weight of an object, that can be detected is a constant proportion of the stimulus level. (i.e. a linear relationship).
These proportions were called Weber Fractions.
Example: Object 1 must weigh 1/40th more/less than Object 2 for the difference to be noticeable or a JND (just-noticeable difference)
Fechner’s law
A principle describing the relationship between stimulus magnitude and resulting sensation magnitude such that the magnitude of subjective sensation increases proportionally to the logarithm of the stimulus intensity
S = k log R
Example:
100 candles is twice as bright as 10.
10 candles is twice as bright as 1.
Stevens’ power law
(1962) is a proposed relationship between stimulus energy and perceived intensity is a power function.
Sensation = a * Intensity b
Chronological Summary of Laws
Weber’s Law:
As stimulus level increases or decreases, the magnitude of change must increase proportionately (linearly) to remain noticeable.
Fechner’s Law:
The magnitude of subjective sensation increases proportionally to the logarithm of the stimulus intensity.
Steven’s Power Law:
Stimulus energy and perceived intensity is a power function.
Doctrine of specific nerve energies:
A doctrine formulated by Johannes Müller (1801–1858) stating that the nature of a sensation depends on which sensory fibers are stimulated, not on how the fibers are stimulated
Photic Sneeze Reflex
Doctrine of specific nerve energies example
Also known as Sun Sneezing
18-35% of the population
2/3 of sun sneezers are female
Correlated with having a nasal septum deviation
Amazingly the cause is still unknown
Caused by light intensity, not spectral composition
Most believe it is caused by “crossed wires”
Icy Hot
Doctrine of specific nerve energies example
Dulls the pain
2 Active Ingredients:
Capsaicin:
Found in chili peppers
Activates warmth fibers
Menthol:
Activates cold fibers
Sensation of both hot and cold
No actual heat transfer
Hermann von Helmholtz (1821–1894)
First person to measure the speed of neural impulses
Demonstrated that neurons follow the laws of physics, Against what Müller believed
Invented the ophthalmoscope
Wrote On the Sensations of Tone, one of the first studies of auditory perception
A ton of other stuff for which there is not room to list.
Speed of neural transmission
~50-100 meters/second
Santiago Ramón y Cajal
(1852–1934)
Discovered the direction of travel of nerve impulses.
Only ~1% of neurons absorb stain
Improved upon a previous (Golgi’s) staining method and drawings.
Created incredibly detailed drawings of neurons and neural structure.
Ramón & Golgi were 1906 Nobel prize co-winners.
Cajal discovered the direction of travel of nerve impulses in the brain and spinal cord.
He was the first to note that information travels one way from the dendrites to the axon and not the reverse.
**He postulated that neurons are discrete entities.
He was unhappy that, because Spanish was not used in the scientific community, his work was not read outside Spain.
Many so-called discoveries by English, German, and French scientists were actually rediscoveries of his work, which had been previously published in Spanish journals.
neurons
The processing of perceptions, thoughts and actions in the brain is accomplished by networks of small cells called neurons.
Dendrites and Soma
Receive information from other neurons
Axon
Conducts that information to other parts of the brain, sometimes over great distances
Axon terminals
Transmit information to other neurons
The Electric Neuron
Molecules in neural tissue are “charged”: + or -
Charged Ions are potassium K+, sodium Na+, chlorine Cl-, and
Proteins-
In the absence of stimulation, the inside of the neuron is slightly more negative than the outside (-70 mv).
This is called the resting potential.
resting potential.
In the absence of stimulation, the inside of the neuron is slightly more negative than the outside (-70 mv).
Ion channels pump ions across the membrane.
The Na+–K+ pump preserves the cell’s resting potential by maintaining a larger concentration of K+ inside the cell and Na+ outside the cell. The pump uses ATP as energy.
synapse
The concentration of ions inside and outside of a neuron can be affected by neurotransmitters that are released into the synapse
Within the axon terminals, the relative charge between the inside and outside of a neuron is called the presynaptic potential.
Within the dendrites and soma, the relative charge between the inside and outside of a neuron is called the postsynaptic potential.
postsynaptic potential.
Within the dendrites and soma, the relative charge between the inside and outside of a neuron.
When neurotransmitters are absorbed by the dendrites or soma, they can alter the postsynaptic potential.
presynaptic potential
Within the axon terminals, the relative charge between the inside and outside of a neuron
Neural Transmission
When neurotransmitters are absorbed by the dendrites or soma, they can alter the postsynaptic potential.
Excitatory neurotransmitters cause the charge to become depolarized (more positive). This change is called an excitatory postsynaptic potential (EPSP).
Inhibitory neurotransmitters cause the charge to become hyperpolarized (more negative). This change is called an inhibitory postsynaptic potential (IPSP).
Both EPSPs and IPSPs are produced by altering the relative concentration of ions between the inside and outside of the cell membrane.
excitatory postsynaptic potential (EPSP).
Excitatory neurotransmitters cause the charge to become depolarized (more positive).
This change is called an excitatory postsynaptic potential (EPSP).
inhibitory postsynaptic potential (IPSP).
Inhibitory neurotransmitters cause the charge to become hyperpolarized (more negative).
This change is called an inhibitory postsynaptic potential (IPSP).
Gated Ion Channels, EPSPs
Causes depolarization of Post-synaptic neuron
Na+ transduction
Gated Ion Channels, IPSPs
Causes hyperpolarization of Post-synaptic neuron
K+ and Cl- transduction
IPSPs and EPSPs are Graded Potentials
They vary in magnitude based on the quantity of neurotransmitters with which they are stimulated.
They travel passively and are attenuated with distance.
At the base of the axon (called the axon hillock), a cell’s potential is determined by the sum of all the excitatory and inhibitory inputs in the dendrites and soma.
action potential.
If the voltage at the axon hillock reaches a threshold of -50 mv, then it will increase rapidly to 50 mv, and then quickly rebound to -70 mv.
This sequence of voltage changes is called an action potential.
During the absolute refractory period it is impossible to generate a new action potential
During the relative refractory period a new action potential can be generated, but it requires a stimulus that is much stronger than usual.
The Toilet Metaphor
The action potential is non-decremental.
That is to say, its magnitude does not change as it travels along the axon.
This is quite different from graded potentials (EPSPs & IPSPs).
Their magnitudes become diminished as they travel within the dendrites & soma, and are therefore referred to as decremental.
When an action potential reaches an axon terminal, it causes neurotransmitters to be released into the synapse.
The stimulation of a neuron modifies the frequency at which it generates action potentials
Excitation and Inhibition
Any neuron can excite or inhibit another neuron based on the type of neurotransmitter it releases from its axon terminals.
The stimulation of a neuron modifies the frequency at which it generates action potentials
What is a wave?
Lights and sounds are composed of waves.
A wave is a type of internal motion of a medium, in which the displaced portion returns to equilibrium.
This disturbance propagates in space as well.
Wave: disturbance propagates in x
Transverse Waves
This is what occurs in the vibrations of stringed instruments
Longitudinal Waves
This is what occurs in the vibrations of wind instruments
Sound Waves
Sound waves involve the longitudinal oscillations of air molecules.
The sound vibrations in a long, narrow tube, such as a trombone, flute or trumpet, propagates in one direction.
In open air, a sound wave propagates radially in all directions
Light Waves
Light waves involve transverse oscillations in electric and magnetic fields
Light
A wave; a stream of photons, tiny particles that each consist of one quantum of energy.
Diagram of a wave
All waves are caused by vibrations
Velocity, Frequency and Wavelength
Within a given medium, light & sound waves travel at a constant velocity.
Thus, long wavelengths oscillate at low frequencies, and short wavelengths oscillate at high frequencies.
Two waves with the same velocity and different wavelengths will have different frequencies.
Two waves with the same frequency and different wavelengths will have different velocities.
Interference
The addition and subtraction of waves.
When two or more waves come together, the individual displacements at each point in space are added together to produce a composite wave.
Absorption
The conversion of energy to a different form, such as heat, when a wave hits an obstacle
When a wave hits a boundary between two media, some of its energy may be converted to heat.
Reflection
The reversal of direction when a wave hits an obstacle
When a wave hits a boundary between two media, some of its energy rebounds in a different direction
Some animals, such as bats, are able to use reflections of sounds to determine the distance of objects
Refraction
The bending of a wave as it crosses the boundary between two media
What happens to a wave when it changes speed
When a wave travels from a less dense to a more dense medium at an angle off the line of normal, it slows down and bends towards the line of normal.
When a wave travels from a more dense to a less dense medium at an angle off the line of normal, it speeds up and bends away from the line of normal.
Refraction is what causes a pencil to appear bent when it is partially submerged in water.
Diffraction
The bending of a wave around an obstacle.
Waves can sometimes bend around obstacles.
Long wavelengths diffract more than short wavelengths
Diffraction through a slit.
Interference patterns can occur when a wave interacts with its own reflection
Wave Interactions
Interference – The addition and subtraction of waves
Absorption – The conversion of energy to a different form, such as heat, when a wave hits an obstacle
Reflection – The reversal of direction when a wave hits an obstacle
Refraction – The bending of a wave as it crosses the boundary between two media
Diffraction – The bending of a wave around an obstacle
Scattered
Energy that is dispersed in an irregular fashion.
When light enters the atmosphere, much of it is absorbed or scattered and never reaches the perceiver.
Transmitted
Energy that is passed on through a surface (when it is neither reflected nor absorbed by the surface)
When a wave hits a boundary between two media, some of its energy may be transmitted across the boundary
Short wave lengths refract more than long wavelengths.
Rainbows are caused by light that is refracted through particles of water.
Rule to remember
Short things have higher natural frequencies than long things
Natural Frequencies
Most objects have a specific frequency at which they vibrate most readily. This is called the natural or fundamental frequency.
For a vibrating string, the natural frequency increases with the tension of a string, and decreases with increasing length or mass.
For a vibrating tube, the natural frequency decreases with increasing tube length, and is higher for open tubes than for closed tubes.
The manipulation of these parameters is what allows musicians to play musical instruments.
Rule to remember: Short things have higher natural frequencies than long things
Tacoma Narrows bridge disaster
Shows what can happen when waves hit an object at its natural frequency.
This is also the reason why singers are able to break a glass by singing at the appropriate pitch.
Methods of representing waves
Waveform plots
Spectral plots
Spectrograms
sine waves
Sound is a change in the pressure of the air.
The waveform of any sound shows how the pressure changes over time.
The eardrum moves in response to changes in pressure.
Any waveform shape can be produced by adding together sine waves of appropriate frequencies, amplitudes and phases.
The amplitudes of the sine waves give the amplitude spectrum of the sound.
The amplitude spectrum of a sine wave is a single point at the frequency of the sine wave.
Spectrum
Amplitude against frequency
Waveform
Amplitude against time
Amplitude
Amplitude is a measure of the pressure change of a sound and is related to how loud the sound is.
Amplitude squared is proportional to the energy or intensity (I) of a sound.
adding sine waves
A sound which has more than one (sine-wave) frequency component is a complex sound.
A periodic sound is one which repeats itself at regular intervals.
A sine wave is a simple periodic sound.
Musical instruments or the voice produce complex periodic sounds.
They have a spectrum consisting of a series of harmonics.
The lowest frequency (of which all the others a re multiples) is called the fundamental frequency.
Each harmonic is a sine wave that has a frequency that is an integer multiple of the fundamental frequency.
The left column shows individual harmonics; the right shows their sum; the yellow panels show the amplitude spectrum of the sound.
Fourier Synthesis
Here is 1/20th of a second of the waveform and also the spectrum of a complex periodic sound consisting of the first four harmonics of a fundamental of 100 Hz.
All the frequency components are integer multiples of 100 Hz.
A periodic sound consists of a section of waveform that repeats itself.
The period of the complex wave is the duration of this section.
In this case it is 1/100s or 0.01s, or 10 ms.
The period is the reciprocal of the fundamental frequency (in this case 100 Hz).
If you change the period of a complex sound, you change its pitch.
Shorter periods - higher fundamental frequency - higher pitch.
Harmonics are integer multiples of the fundamental frequency
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A spectrogram is a 3-dimensional plot of frequency and amplitude as a function of time.
Amplitude is represented in a spectrogram by shades of gray
Spectrograms are especially useful for representing complex waveforms like speech that change over time
Properties of Waves
Amplitude:
Controlled by the magnitude of the forces that started the wave
Frequency f of oscillations:
Controlled by forces starting the wave and by the nature of the material or object that is oscillating
Spectral Composition:
Refers to the mixture of different frequencies in a wave
The 3 Main Perceptual Attributes of Sound
Loudness (is most related to intensity)
Pitch (is most related to frequency)
Timbre (is most related to spectral composition)
The terms pitch, loudness, and timbre refer not to the physical characteristics of sound, but to the mental experiences that occur in the minds of listeners.
The 3 Main Perceptual Attributes of Light
Brightness (is most related to amplitude)
Hue (is most related to wave length or frequency)
Saturation (is most related to spectral composition)
The terms Brightness, Hue, and Saturation refer not to the physical characteristics of light, but to the mental experiences that occur in the minds of observers.
The stimulus for vision is …..
visible electromagnetic radiation, which can be be characterized by its wavelength.
The human visual system is sensitive to wavelengths from 400 to 700 nanometers (10-9 meter)
The human eye is made up of various parts:
Cornea: The transparent “window” into the eyeball
Aqueous humor: The watery fluid in the anterior chamber
Pupil: The dark circular opening at the center of the iris in the eye, where light enters the eye
Crystalline lens: The lens inside the eye, which enables changing focus. Focus is controlled by ciliary muscle.
Zonules of Zinn: connect the ciliary muscles with the lens
Vitreous humor: The transparent fluid that fills the vitreous chamber in the posterior part of the eye
Retina: A light-sensitive membrane in the back of the eye that contains rods and cones, which receive an image from the lens and send it to the brain through the optic nerve.
Cornea
The transparent “window” into the eyeball
no blood supply, but has nerves to feel scratches and dryness. highly organized structure to let light through
MOST OF THE REFRACTION HAPPENS HERE!!
Quick regeneration.
Contact lenses: sit on a layer of tears in front of cornea.
Aqueous humor
The watery fluid in the anterior chamber
nutrient & oxygen delivery
Pupil
The dark circular opening at the center of the iris in the eye, where light enters the eye
Crystalline lens
The lens inside the eye, which enables changing focus.
Focus is controlled by ciliary muscle.
Zonules of Zinn
connect the ciliary muscles with the lens
Vitreous humor
The transparent fluid that fills the vitreous chamber in the posterior part of the eye
FLOATERS
bio-debris.
No concern.
Egg-whites!!
Retina
A light-sensitive membrane in the back of the eye that contains rods and cones, which receive an image from the lens and send it to the brain through the optic nerve.
Refraction is necessary to focus light rays and this is done by the cornea and the lens
The lens can change its shape, and thus alter the refractive power: Accommodation !!!
TAPETUM
the colorful, shiny material located behind the retina that reflects light back through the retina to get a second chance at capturing missed photons!
A layer of tissue in the eye of many vertebrate animals.
It lies immediately behind the retina.
It reflects visible light back through the retina, increasing the light available to the photoreceptors.
This improves vision in low-light conditions, but can cause the perceived image to be blurry from the interference of the reflected light
Cataracts
loss of transparency in lens (solved with silicone implants)
Presbyopia
“old sight”
Inability to accommodate nearby objects.
Pigment Epithelium
nourishes receptors, non-reflective in humans & absorbs stray light
Receptors face away from light.
Photoreceptors are in the LAST LAYER, mostly because of the pigment epithelium (provide vital nutrients to the photoreceptors
Photoreceptors
Cells in the retina that initially transduce light energy into neural energy.
100 million photoreceptors
Chemical signals.
Rods
Photoreceptors that are specialized for night vision
Cones
Photoreceptors that are specialized for daylight vision, fine visual acuity and color
CHROMOPHORE
the light-catching part of the visual pigments of the retina.
4 different types of pigments.
Cones have three different kinds which respond to Long, medium and short wavelengths.
Visual process
biochemical cascade of events:
1. Closing of channels to the outersegment.
2. This causes HYPERPOLARIZATION in the cell body.
3. Reduction of neurotransmitter (GLUTAMATE) at the synaptic level.
Bipolar cell knows a photon has been caught.
GRADED POTENTIALS (not action potentials): the more photons, the less neurotransmitter.
5 million cones
120 million rods
2 million ganglion cells
1 degree of visual angle where no RODS, only cones. Right behind center of pupil.
Therefore, many receptors must send signals to each ganglion cell an this is called convergence.
The extent of convergence on different parts of the retina produces a tradeoff between sensitivity to low light levels and sensitivity to fine spatial detail.
The retina’s horizontal pathway:
Horizontal and amacrine cells
Horizontal cells: Specialized retinal cells that run perpendicular to the photoreceptors and make contact with photoreceptors and bipolar cells
Responsible for lateral inhibition, which creates the center–surround receptive field structure of retinal ganglion cells
Amacrine cells: These cells synapse horizontally between bipolar cells and retinal ganglion cells
Have been implicated in contrast enhancement and temporal sensitivity (detecting light patterns that change over time)
Horizontal cells
Specialized retinal cells that run perpendicular to the photoreceptors and make contact with photoreceptors and bipolar cells
Responsible for lateral inhibition, which creates the center–surround receptive field structure of retinal ganglion cells
Amacrine cells
These cells synapse horizontally between bipolar cells and retinal ganglion cells
Have been implicated in contrast enhancement and temporal sensitivity (detecting light patterns that change over time)
The retina’s vertical pathway:
Photoreceptors, bipolar cells, and ganglion cells
Bipolar cell: A retinal cell that synapses with one or more rods or cones (not both) and with horizontal cells, and then passes the signals on to ganglion cells
Diffuse bipolar cell: A bipolar cell that receives input from multiple photoreceptors
Midget bipolar cell: A small bipolar cell that receives input from a single cone
P ganglion cells
Connect to the parvocellular pathway
Receive input from midget bipolar cells
Parvocellular (“small cell”) pathway is involved in fine visual acuity, color, and shape processing.
Poor temporal resolution but good spatial resolution.
M ganglion cells:
Connect to the magnocellular pathway
Receive input from diffuse bipolar cells
Magnocellular (“large cell”) pathway is involved in motion processing.
Excellent temporal resolution but poor spatial resolution
Ganglion Cells
last stage before information leaves the eye and travels to and through the brain!
Rod vision more sensitive to light than cone vision.
There is greater convergence of rods than cones onto ganglion cells
Thus, there is greater summation of rod signals,
and less stimulation per rod is required to obtain a response.
Cone vision can see finer details than rod vision.
Greater convergence of rods than cones onto ganglion cells limits their spatial sensitivity.
ON-center ganglion cells
Excited by light falling on center, inhibited by light falling surround
OFF-center ganglion cells
Inhibited by light falling on center, excited by light falling on surround
Ganglion cells that receive inputs from the fovea have smaller receptive fields than cells that receive inputs from more peripheral regions.
For an on centered ganglion cell, the response rate is greatest when the stimulus just fills the excitatory central region.
When the stimulus covers the entire receptive field, the cell will fire at its background rate.
Receptive field
The region on the retina in which stimuli influence a neuron’s firing rate.
Ganglion cells are unaffected by average light intensity, and are most responsive to DIFFERENCES in light intensity.
Hermann Grid
Why do spots appear at the junctions, and why do they disappear when a junction is fixated?
Hermann Grid
Why do spots appear at the junctions, and why do they disappear when a junction is fixated?
Spots do not appear at the fixated junctions because receptive fields in the fovea are smaller than in more peripheral regions.
Cells with receptive fields at the junction receive more inhibition (two more minus signs), so that the junctions appear darker.