Prelim 1 Flashcards
Fossil
remains, traces, or impressions of once living organisms ie. skeleton, impression, cast, trace, or coprolite (poop); most found in sedimentary rocks
Forces that impede fossilization
natural processes such as predators/scavengers, bacterial decay (soft tissue), dissolution in water (soft and hard tissue), or physical disturbance (wave action, wind)
Conditions that promote fossilization
rapid burial, protection from physical disturbance ie. quiet, deep water, anaerobic conditions (prevent bacterial decay)
Sedimentary rock formation
formed from the deposition of sediment falling to the bottom of a body of water
What gets preserved in fossils of animals
hard parts ie. teeth, bone, chitinous exoskeleton, or calcium carbonate shells
What gets preserved in fossils of plants
seeds, pollen, leaves, wood, rarely flowers (fragile petals)
What gets preserved in fossils of microbes
bacteria, microbial mats ie. stromatalites are formed from biofilms of cyanobacteria that trap sediment which eventually harden and form layers
Types of preservation
original remains (skeletal body, other body elements), permineralization/petrification, trace fossils, impression fossils, casts and molds
Permineralization/petrification
process where minerals are deposited in tiny holes within bones, or wood and over time completely replacing the original organism and all that remains is a stone structure
Impression fossils
made up of carbonaceous film imprint of an organism
Lagerstatten
German word that means “storage place”, place where fossils are exceptionally preserved (numerous and well preserved)
Burgess Shale
a famous Lagerstatten from the Cambrian which has yielded many of the organisms that contribute to our understanding of the Cambrian explosion
How do we know the age of a fossil?
relative and absolute dating
Geological chronology
the science of dating geological layers and fossils
4 principles relative dating is based on
superposition, original horizontality, lateral continuity, and cross-cutting
Principle of superposition
geological layers are formed by laying one on top of the other so that the youngest layer is on top
Principle of original horizontality
layers are first deposited horizontally and then they may be deformed later such as from the movement of continental plates
Principle of lateral continuity
layers continue laterally over distances; each layer is deposited at the same so that even if erosion has removed some of the layer, the layer is still the same layer after the gap
Principle of cross-cutting
if there is a cross-cutting layer or intrusion in rock layers, the intrusion is always younger than any of the layers it is cross-cutting
Index fossils
fossil organisms that are only found in a particular rock layer and are also geographically widespread so that the layers can be stratiagraphically correlated with each other in different locations according to the principle of lateral continuity
How was geological time scale (GTS) created?
by correlating layers based on index fossils; derived from the spatial distribution of rocks and the vertical sequence of rocks and contained fossils
How/when did absolute dating become possible
with the discovery of radioactivity in the late 19th century
Absolute dating
done by examining the radioactive decay of unstable isotopes
Radioactive decay
decay of a parent isotope gives rise to a stable daughter isotope at some characteristic rate of decay
Which rocks can be absolute dated
only igneous rocks; clock starts ticking when rock solidifies (daughter isotope is 0 in molten rock); sedimentary rock decays too quickly (carbon decays too quickly)
Radiometric dating: half lives
0 half lives = 100% parent isotope; 1 half life = 50%; 2 half lives = 25%; 3 half lives = 12.5% etc.
Uranium-lead effective dating range
10 million - 4.6 billion years
Potassium-argon effective dating range
100,000 - 4.6 billion years
Carbon-14 effective dating range
100 - 100,000 years
Phylogeny
a visual representation of the evolutionary history of populations, genes, and species
Tips of phylogenetic tree
represent groups of descendant taxa; most often species but can also represent molecules or populations
Branches of phylogenetic tree
lineages evolving through time between successive speciation events
Node of a phylogenetic tree
a point in a phylogeny where a lineage splits (a speciation event)
Clade/monophyletic group
an organism and all of its descendants; consists of the most recent common ancestor and all of its descendants
Paraphyletic clade
an ancestor and a group of taxa but it is not monophyletic because some of the descendants are missing
“Tree Thinking”
using data to construct trees, and reading trees to determine evolutionary relationships
Pedigree vs Phylogeny
pedigree: individuals, 2 ancestors, unlimited descendants; phylogeny: populations, 1 ancestor, 2 descendants (except for a polytomy)
How do we infer relatedness in a phylogeny
based on sharing of derived characteristics
Character
anything inherited (genetically determined, or DNA sequence itself) that can be used to determine relationships; morphological, physiological, (traits) or molecular (DNA sequence)
Ancestral state
the historical state of a character
Derived state
the more recently evolved state of a character
Synapomorphies
shared derived traits which are phylogenetically informative
Polytomy
describes an internal node of a phylogeny with more than 2 branches (the order in which the branching occurred is not resolved)
Choosing optimal phylogeny
select tree with the smallest number of character state changes (most common for morphological characters); select tree that is the most probably (based on probability methods, typically used for DNA mutations)
What are phylogenies used for?
map characters, trace the origins of epidemics, or to inform taxonomy; is it NOT a depiction of the degree of similarity
Bootstrap/posterior probability as branch support
statistical confidence assigned to certain branches
Neutral theory of molecular evolution
at the fine scale level, most new mutations are not favored or disfavored by natural selection (synonymous mutations); neutral mutations will arise at random and random processes (ie. genetic drift) will determine their fate in a population; since mutations arise at an average rate, they can be used to date the nodes on a molecular phylogeny
Graphic variations of a phylogeny
Cladogram (branching only); Phylogram (degree of change); Chronogram (calibrated to real time)
Parsimony inference
the best phylogeny is the one that explains the observed character data by positing the fewest evolutionary changes
Homology
shared traits because they are inherited from a common ancestor
Autapomorphy
a trait that does not help us distinguish between two trees because it is only in one lineage
Homoplasy
when similar characteristics are not due to common ancestry but instead arise by convergent evolution or evolutionary reversals; can create the mistaken impression that two species are closely related when they are not
Vestigial
a trait that has become functionless in the course of evolution (but still is present)
Ideal characters for phylogenies
have low rates of evolutionary convergence and/or reversal
What do different speeds of evolving characters show ie. fast vs slow)
slowly evolving characters ( including DNA sequences) can show the relationship between distant taxa while rapidly evolving characters (including DNA sequences) can reveal relationships between closely related taxa
Index case
source of the outbreak ex. someone who is initially affected brings a human pathogen to a new geographic location and transmits the disease to a few other recipients