Plant systems - Xylem Flashcards
Xylem
Transports water and dissolved mineral ions
Vascular tissue
In roots:
-Xylem is central and star shaped
-Phloem in between the groups of xylem cells
In stems:
- Vascular bundle in a ring
- Phloem outer side
- Xylem inner side
In leaves:
-Vascular tissue in the midrib and in a network of veins
Root structure
- Root hair
- Epidermis (Epi = outer)
- Cortex between epidermis and endodermis
- Endodermis (Endo = inner)
- Star shaped xylem
- Phloem in space between xylem cells
Root tissue description
Epidermis:
- Outer layer of small thin-walled cells with no waxy cuticle
- Root hair cells grow outwards to form root hairs
- Root hairs have large surface area for the uptake of mineral ions, oxygen and water
Cortex:
- Made up of unspecialised parenchyma cells which may store starch grains
- Large spaces between parenchyma cells makes it easier for oxygen to diffuse into the root
Endodermis:
- Single layer of cells which forms a ring around the vascular bundle
- Cell walls of endodermal cells contain waxy waterproof suberin
- Suberin forms the Casparian strip which influences the route by which water enters the vascular bundle from the cortex
Pericycle:
- Layer of lignified sclerenchyma cells
- Provides mechanical support
Absorption of water
- Water potential inside the epidermal cells is lower than surrounding soil due to high concentration of solutes
- Water is absorbed into the epidermal cells of the root by osmosis
Absorption of mineral ions
- Cytoplasm of epidermal cells generally have a much higher solute concentration than the surrounding soil
- Ions are actively transported into the epidermal cells against the concentration gradient, moving along the apoplast pathway until they reach the endodermis
- The Casparian strip blocks the apoplast pathway so ions enter the xylem by diffusion or active transport
- The blocking of the apoplast pathway then allows for ions to be selectively taken up by the endodermis
Pathways of water across the cortex
Water can follow 3 routes across the cortex.
Apoplast pathway:
- The cellulose cell walls are freely permeable so water passes freely from one cell to another
- Water is pulled across the cortex due to forces of cohesion between water molecules
- This is the fastest and most significant pathway
Symplast pathway:
- Water diffuses down its concentration gradient through the cytoplasm of adjacent cells
- Strands of cytoplasm called plasmodesmata pass through pores in the cell walls, forming a continuous pathway for water movement
Vacuolar pathway:
- Water diffuses down the same concentration gradient as in the symplast pathway
- Water moves from the cytoplasm and across the tonoplast of the vacuole and back out into the cytoplasm
Pathway of water from the endodermis to the xylem
- The Casparian strip blocks the apoplast pathway due to the waterproof substance suberin
- Water is forced into the symplast pathway
- Water passes through the pericycle and into the xylem vessels via the symplast pathway down a water potential gradient
Stem Structure
- Epidermis
- Cortex
- Phloem
- Cambium
- Xylem
- Pith
Stem tissue description
Epidermis:
- Single layer of cells
- Helps maintain the shape of stem
Cortex:
-Made up of chlorenchyma cells which have extra cellulose in the corners for added mechanical support
Vascular bundle:
- Arranged in a ring pattern
- Gives the stem strength and flexibility
Cambium:
-Undifferentiated tissue which has the ability to differentiate into either phloem or xylem
Pith:
- The centre of the stem
- Consists of parenchyma cells
Xylem vessel elements
Empty cells joined end to end, lined with lignin, only found in angiosperms (flowering plants)
Formation:
- Begin as normal plant cells arranged end to end
- As the xylem develops, a hard waterproof substance call lignin is laid down within the cell walls
- Lignin is laid down initially in young plants in a circular pattern which allows the young plant to grow
- As the plant matures the cell dies, and the spirals partially join up, providing strength and support
Function:
-Xylem vessels have pits which allow the sideways movement of ions and water into adjacent xylem vessels
Tracheids
- Found in primitive plants
- Like xylem they are dead cells
- Their cell walls also contain lignin and pits
- They are elongated spindle shaped cells with tapering ends
- Water takes a twisting path up the plants
Movement of water from root to leaf
Water movement involves ALL 3 principles:
-Root pressure (Push)
- Cohesion-tension theory (Pull)
- Transpiration stream (Pull)
Root pressure
- Active transport of solutes in the root lower the water potential in the xylem
- The influx of water by osmosis raises the hydrostatic pressure
- This creates a positive root pressure and pushes water up the xylem
Cohesion tension theory
- Capillary action is the movement of water up narrow tubes
- It occurs due to forces of cohesion between water molecules and adhesion between the lining of the tube
- It is a passive process which can occur in dead tissue