Plant Hormones and Sensory Systems Flashcards
Hormones in Plants
- Hormones travel throughout the body via the vascular tissues (xylem + phloem) and cell-to-cell (plasmodesmata)
- potentially every plant cell can produce plant hormones
- plants DO NOT have specialized hormone-producing glands like animals
Hormone Roles in Plants
Hormones regulate a variety of plant behaviors to different stimuli/environmental conditions; hormones:
1. initiate and regulate plant behaviors
2. provoke these responses and pathways that regulate the responses
Auxin
the “youth” hormone and master growth regulator
Tissues with high auxin are areas of new growth and do not exhibit signs of aging
- Responsible for cell elongation in phototropism + gravitropism, apical dominance, and inhibition of abscission
- synthetic auxin is used as a rooting hormone to promote root growth on cuttings + detached leaves
Senescence
ageing
Abscission
lead falling in plants
Cytokinin
the “cell division” hormone
- promotes CYTOKINESIS (cell division)
- abundant in growing tissues, such as roots, embryos, and fruits, where cell division is occurring
- INFLUENCED BY AUXIN (EX: apical dominance results from a balance of auxins that inhibit lateral buds and cytokinin that promote bushier growth)
Gibberellin (GAs)
the “growth” hormones; stem, fruit, and seed growth
- Stimulate shoot elongation, seed germination, and fruit + flower maturation
- synthesized in the root and stem apical meristems, young leaves, and seed embryos
- breaks dormancy in the seeds of plants that require exposure to cold or light to germinate
Abscisic Acid (ABA)
the “dormancy” hormone
- accumulates as a response to stressful environmental conditions (dehydration, cold temps, or short day lengths)
- its activity COUNTERACTS THE GROWTH PROMOTING EFFECT OF GAS AND AUXINS
- causes abscission of leaves, inhibits stem elongation, induces dormancy in buds and seeds, and closes stomata in short-term droughts
Ethylene
the “aging” hormone; lead abscission, flower wilting, and lead abscission
- aging tissues and nodes of stems produce ethylene
Plant Responses to Stimuli
- Light: Phototropism + Germination
- Gravity: Gravitropism
- Growth
- Water or Water Stress
- Touch: Thigmotropism
Plant Response to Light
- Plants can detect and respond to BLUE, RED, and FAR-RED light
- Different wavelengths are detected by different photoreceptors (protein + Chromophore = CHROMOPROTEIN)
- Behaviors regulated by light include phototropism, stem elongation, germination, and photoperiodism
Phototropism
movement toward/away from light; movement in response to light
- light is detected by the tip of the plant (apical meristem)
- the response occurs when the signal travels from the apical meristem to the base of the plant causing BENDING
- the detection of light in the apical meristem occurs via PHOTOROPINS (phot1 + phot2), which detect BLUE LIGHT
- Auxin promotes elongation, causing bending
Acid Growth Hypothesis
Auxin stimulates cell elongation of the shady side of the stem through the ACID GROWTH HYPOTHESIS:
- auxin causes cells to activate proton pumps, which pumps protons out of the cells and intro the space between the PM and cell wall
the movement of protons into the extracellular space causes:
1. lower pH activates EXPANSIN
2. Sugar movement
Phototropism: Expansin
the lower pH activates this, which breaks the links between the cellulose fibers in the cell walls, making them more flexible
Phototropism: Sugar Movement
the high concentration of positive ions causes sugars to move into the cell, which creates an osmotic gradient where water moves into the cell causing it to expand
Phototropism Response Summary
- Phototropins Phot1+Phot2 are present in the plant apical meristem
- When activated by BLUE light, phot 1+2 cause the accumulation of auxin on the shaded side of the plant
Auxin promotes elongation due to the weakening of the cell wall combined with an influx of water that stretches the cell - since the cell expansion only occurs on the shaded side of the stem, the plant bends away from the shade and toward the light
Response to RED light: Growth/Germination
- RED LIGHT PROMOTES STEM ELONGATION; indicates full sun to a plant
- A far-red light indicates that a plant is being shaded out by another plant
- unfiltered, full sunlight conditions contain much more red light than far red light
- PLANTS USE RED VS FAR-RED LIGHT DETECTINO TO GROW AWAY FROM SHADE AND TOWARDS LIGHT
Phytochrome
the chemo protein responsible for red/far-red light detection; acts like a reversible switch that can absorb either red or far-red light
- Pr (phytochrome red)
- Pfr (phytochrome far-red)
Pr
capable of absorbing red light (~667 nm)
- phytochrome is in this form when in the dark, causing a seed to not germinate
Pfr
capable of absorbing far-red light (~730 nm)
- the physiologically active form of the protein; because phytochrome is in the Pfr state after exposure to red light, this means that exposure to red light turns the phytochrome “on”
- in dormancy + germination, this signaling causes the trancription of AMYLASE
Pr + Pfr
- when Pr absorbs red light, it is immediately converted to Pfr; when Pfr absorbs far-red light, it is immediately converted back to Pr
- Exposure the far-red light inhibits phytochrome activity
Phytochrome: Plant Growth
- phytochrome stimulates plant growth TOWARD RED LIGHT in combination with the hormones cytokinin and gibberellin
- cytokinin is only capable of promoting division when auxin is also present, which is only at APICAL MERISTEMS
- auxin regulates GA levels
Phytochrome: Seed Germination
- the seeds of many plants go into dormancy after fertilization to ensure survival - many different signals can trigger germination depending on plant species
- for many species, this signal is a RED LIGHT, as it provides a signal that the seed is in a good location for cull access to the sun after germination
- a seed that germinates in shaded areas or too deep underground is likely to die after germination
- some plant species initiate germination through a light-independent process regulated by GA
Amylase
breaks down starches stored in a seed into simple sugars, then germination proceeds