Physiology of Olfaction and Gustation Flashcards
two senses for perception of chemicals - what are they?
olfaction (smell)
gustation (taste)
olfaction?
can detect very small amounts of many substances from a great distance
gustation?
can detect larger amounts of relatively few substances in direct contact
functions of olfaction?
detection of food, enemies, navigate terrain/detect territories and detection of potential mates
olfaction developed early in vertebrate evolution - why?
required most complex processing in the brain
mammalian neocortex developed from?
from olfactory sense and a large portion of the neocortex is still devoted to olfactory processing in modern mammals
nostril have slits laterally to?
to direct airflow laterally to prevent interference with incoming smells - stereo nostrils allow directional smelling
what do turbinates do?
they direct airflow - ventrally in normal breathing, dorsally when sniffing
lamina transversa?
(absent in microsmatic species) is a bone that separates the olfactory recess from the main airway and traps scent molecules to enhance detection
olfactory recess?
contains the ethmo-turbinates lined with olfactory epithelium containing all the olfactory receptors (3 x10(8))
cribiform plate?
separates cranial cavity from nasal cavity - a perforated plate to allow olfactory cell processes to pass through and synapse with olfactory bulb mitral cells
olfactory bulb?
processes olfactory information - 40 times greater percentage of brain devoted to olfaction in the dog vs. human
what does the nasal cavity contain?
contains scroll-like bones called turbinates
describe the turbinates in the rostral cavity:
these are highly vascularised and used for heat exchange
describe the turbinates in the caudal nasal cavity?
these are covered in olfactory epithelium
through which three main passage/meatuses can airflow be directed through?
either:
ventral meatus
middle meatus
dorsal meatus
ventral meatus?
shortcut beneath the heat exchanger directly to the nasopharynx a.
middle meatus?
leads air through the heat exchanger and to the caudal part of the olfactory epithelium before entering the nasopharynx
dorsal meatus?
leads air above the heat exchanger and to the olfactory epithelium and then to the nasopharynx
breathing at rest - airflow?
air flows mainly through the two more ventral meatuses
what happens during sniffing?
air passes dorsal to the heat exchanger
the airflow through the nasal cavity becomes turbulent, which further increases the air that reaches the olfactory epithelium
where does exhalation occur?
occurs mainly through the ventral meatus
olfactory receptors respond only to?
only to substances in contact with olfactory epithelium and need to be dissolved in mucus
olfactory adaptation?
over time, olfactory threshold increases so that more of a substance is required to elicit the same response
adaptation/habituation of smell?
smell of odoriferous substance first detected much more noticeable
gradually becomes less noticeable
what is olfactory adaption mediated by?
by Ca2+ ions
degree of adaptation is measured by?
measured by rise in threshold concentration to activate olfactory receptors
what does degree of adaptation vary depending on?
depending on the substance
nasal pain fibres - what nerve?
trigeminal nerve (cranial nerve V)
how many branches of trigeminal nerve for nasal pain fibres?
mandibular branch
maxillary branch
ocular branch
many trigeminal pain fibres from the maxillary branch are found where?
found in olfactory membrane
what are trigeminal pain fibres stimulated by?
by irritating substances
what are trigeminal pain fibres responsible for?
for initiating sneezing, lacrimation and other reflex responses
what is the vomeronasal organ?
it is a sensory organ located in the rostral palatine bone with similar functionality to the olfactory epithelium
vomeronasal organ found in which animals?
early evolutionary organ found in amphibians, reptiles and mammals
absent in fish and lost in birds
vomeronasal organ in mammals?
well-developed and functional in most mammals (latest evidence suggests present but rudimentary in humans)