oceanography Flashcards
origin/meaning of oceanography
okeanos - Oceanus
graphia - recording/ describing
why oceanography isn’t really a correct term
oceanology = study of oceans
oceanography as a pure science
not. it is a compilation of biology, chemistry, physics, geology.
3 broad stages of ocean exploration
1,2 Early investigations
3. Modern investigations
Early investigations focused on
exploring oceans
exploring landmasses
early scientific investigation of ocean
Early oceanography, the explorers
James Cook
Robert Fitzroy
Wyville Thomson
Fridtjof Nansen
Early oceanography, the time
1700-1900 CE
James Cook
1768-1779
3 major voyages
mapped NZ and Aust
data: geo, bio, currents, tides, temps
Robert Fitzroy and Charles Darwin
1831-1836
HMS Beagle
South America, Galapagos
Came out of the HMS Beagles expedition
Two major ecological theories
- Atoll formation
- Natural selection
Wyville Thomson
1872-1876
Circled globe
Explore abyss
data: water chem, temp, currents, biota, sedimentary
Major Thomson discoveries
Refuted abiotic abyss theory
Recorded 7000+ species down to 9km
First sea-bottom topography charts
Abiotic abyss theory
Forbes
no species in abyss
Nansen
1893-1896
circulation of Arctic ocean
Drifted in boat (Fram) for three years locked in sea ice off Siberia, about 2km/yr
Nansen discoveries
no polar continent
water depths along path
water-mass structure
circulation patterns
Modern Oceanography
1900+ CE
Marine institutes
German scientists
Collaborations
Marine institutes
beginning of educating people in oceanography
Scripps, 1903, California
Woods Hole, 1930, Mass.
German scientists
1925-1927
Survey S Atlantic
Echo sounding
Vertical profiles
Major collaborations
International geophysical year, 1957-1958
International Indian ocean expedition, 1959-1965
Deep sea drilling project, 1968-1975
Major discoveries of deep sea drilling project
seafloor spreading
plate tectonics
Current/ future research
- International efforts (cost)
- Technology
Oceanography technology
Submersibles (Alvin)
ROV (Jason)
Computers (Modelling)
Satellites (GPS)
Earths interior, sections
Crust
Mantle
Outer core
Inner core
Earth’s crust
solid
35-50km, 0.4%
Al, Si, O
Earth’s Mantle
Solid / plastic
2900 km, 68%
Mg, Fe, Si, O
Outer core
Molten
2200km thick
inner core
Solid
1300km
Fe, Ni
Earths divisions based on physical characteristics
Lithosphere Asthenosphere Mesosphere Outer core Inner core
Lithosphere
Rigid and brittle
Crust + upper mantle
Asthenosphere
Plastic
intermediate mantle
T>P
Mesosphere
rigid
lower mantle
P>T
Outer core physical state
molten Fe-Ni alloys
T>P
Inner core physical state
solid Fe-Ni alloys
P>T
Earths spheres
Hydrosphere
Atmosphere
Biosphere
Hydrosphere
all ‘free’ water
97% in ocean
10% of total water
Atmosphere
gases
N 79%, O 16%
Remaining 90% of water, not in hydrosphere
locked in rocks
Biosphere
living and non-living
thin but dynamic
organic - C, H, O
Measurement of seafloor topography based on depth
bathymetry
greatly improved after WWII
physiographic provinces
Continental margins
deep ocean basins
midocean ridges
Parts of continental margin
continental shelf
continental slope
continental rise
Continental shelf
up to 1000km wide
0.5 deg slope
ends at 130-200m depth
continental slope
2-3km deep
4 deg slope
steep, v-shape canyons
continental rise
up to 500km wide
1 deg slope
base up to 4km deep
Deep ocean basin
beyond margin
several bathymetric features
bathymetric features
Abyssal plains
abyssal hills
seamounts
deep-sea trenches
Abyssal plain
3-5km deep
100-1000m thick
<0.5 deg slope
Abyssal hill
domes
<1000m tall
100km wide
volcanic rock
Seamount
> 1000m tall
Extinct or active volcano
flat-topped seamount
guyot
deep-sea trench
3-5km deeper than surrounding
against contin. margin
partially sed. filled
steep-sided
Midocean ridges
Connected, >60,000km cover 1/3 of ocean floor mountain ranges rift valley geologically active volcanoes, earthquakes transform faults
max ocean depth
11km
earthquake epicentres
midocean ridges
transform faults
deep sea trenches
earthquake types
shallow and weak
shallow-to-deep and strong
band of earthquakes in subduction zone
benioff zone
benioff zone
45 degrees into earth
subducting plate and melting
after subducting plate melts
molten portion lower density, rises to surface, volcanic arc
Predominant subduction zones
Pacific (ring of fire)
15-45 cm/yr subduction
ocean-ocean convergence produces
andesite
density btw basalt and granite
lithosphere contains
brittle outer shell
crust
upper mantle
3 types of plate boundaries
tension
compression
sliding
sliding plate boundaries
transform faults
tension plate boundaries
divergent zones
driving force of plate tectonics
thermal convection
thermal convection
heat transfer by fluid motion
heat - lower density - rise - convect
currents- draging of plates
cold edge of subducting
slab-pull
subducting plate pulls plate down
water molecule
dipole
bent
105 deg between H
covalent bonds
covalent bonds
share electrons
H2O residual charge
+ at H end
- at O end
most common elements dissolved in seawater
Na+
Cl-
water clusters
irregular grouping of molecules
size decreases w/ increased T
H bonded
Ice density
8% less than water
Ice
open hexagonal crystals
angle btw H expands to 109.5
chemical bonding
water density
max at 3.98 deg C
solutes in seawater
salt ions nutrients gases dissolved metals org compounds
salt ions
major constituents
85.6% Na and Cl
99% w/ sulfate, Mg, Ca, K
particles that don’t change concentration over large areas on average
conservative
salinity
g/ kg seawater
ppt
principle of constant proportion
relative proportions of major constituents are constant
use of principle of constant proportion
can determine S by measuring only one ion
measuring salinity
conventionally measure Cl- and use principle of constant proportion
chlorinity /salinity relation
S = 1.80566 x cholorinity
why measure Cl -
halogen
less reactive
sw freezing pt
-1.91 @ 35 ppt
sw density
greater than fw
adding solutes increases atomic mass
sw vapor pressure
lower than fw
salinity lowers vp
fw evaporates at higher rate
why does salinity lower vp
more molecular bonds
isotherms
parallel to latitudes
vary seasonally
characteristic of tropical, temperate oceans
thermocline
thermocline depths
200-1000m
temperate ocean thermocline
~ inexistent in March
grows during spr-summ
weakens in winter
global salinity
highest btw 20-30 deg
decreases twd poles, equator
surface salinity
- dependent on evaporation and precipitation
- closely follow evap-precip line
polar SST
low
evap and precip both minimal
temperate SST
low
evap moderate
precip max
haloclines occur
40 deg N - 40 deg S
subtropical SST
max
evap max
precip min
tropical SST
medium
evap max
precip max
density =
mass per unit volume
g/cm3
density depends on
temperature
salinity
pressure
pycnocline layer
corresponds with thermocline and/or halocline
outside of pycnocline
surface layer 2% , 100m thick, seasonal
deep layer 80%
tropical pycnocline
corresponds to permanent thermocline
temperate pycnocline
coincides with halocline
primary regulator of gas [ ] in sw
biotic activity
photosyn, resp., decomp
O2 profile
highest at surface
O min zone
Increases then levels in deep
O2 min zone depth
150 - 1500m
pH =
-log10[H+]
what does pH 7 mean
neutral
1/10 million molecules (10^-7) molecules dissociate into H and OH ions
addition of CO2 to SW
lowers ph of water
H2O+ CO2
- > H2CO3
- > H + HCO3 -
- > CO3 + 2H
carbonic acid
H2CO3
pH of normal sw
7.8 - 8.2
most inorganic C is in the form
bicarbonate
89%
HCO3
main source of dissolved ions
rivers
solar energy
stratifies water column
photosynthesis
air constituents
N 78
O 21
CO2 + halogens + water vapor +… 1%
Pressure =
pgh
p=ro= density
coriolis
In NH deflection to right
coriolis becuase
velocity of rotation different at poles relative to equator
strength of coriolis dependent on
speed
location
equatorial air currents
divergent
heat - rising air - high pressure - circulates
polar air currents
low pressure - cooled air - sinking - convergence
atmospheric cells
hadley - equator
ferrel - mid lats
polar
atmospheric air movements
Northeast trades, Hadley
Westerlies, Ferrel
Polar easterlies, polar cell
wind-driven current
from frictional drag
4% of wind speed
midlatitude currents
flows eastward from the westerlies
low lat currents
flow westward from trade winds
Ekman transport
net flow of water to the right (NH) of the wind 45 deg - drags the layer below - that layer moves 45 deg to the right - drags next layer - .. etc
Depth that Ekman transport effects
100-200m
net water transport due to Ekman
to the right 90 deg
Water movement on east side of continents (NH)
deflected to the right away from continent, deeper water moves up to replace, upwelling
NH gyres
water deflected to the right all the way around the gyre - convergence in middle - downwelling
series of parallel, counter-rotating circulation cells
langmuir circulation
langmuir circulation direction
long axis aligned parallel to wind
Langmuir characteristics
- wind >= 3.5 m/s
- 10-50 m wide
- 5-6 m deep
- several km long
thermohaline upwells where
pacific and indian
classification of organisms (lifestyle)
plankton
nekton
benthos
forms of plankton
phytoplankton
zooplankton
bacterioplankton
virioplankton
forms of benthic organisms
epifauna
infauna
epiflora
classification of organisms (size)
megaplankton (jellies) macroplankton (krill) mesoplankton (copepod, foramin.) microplankton (coccolith.) nanoplankton (diatoms, dinof.) picoplankton (bacteria) femtoplankton (viruses)
classification of organisms (life-history)
Holoplankton
Meroplankton
organisms which are planktic their whole lives
holoplankton
distribution of marine species closely follows
isotherms
rates of biological activity
double per 10 deg. rise
polar organisms
grow slower
reproduce less
live longer
physical process where molecules move from areas of high [ ] to low [ ]
diffusion
pressure depth relationship
1 atm per 10 m
marine fish osmoregulation
body fluid less saline than water
- osmotic water loss
- low urine prod.
- drink SW
- excrete salt through gills
terrestrial food chains
ca. 3 links
diffusion of water molecules through a semipermeable membrane
osmosis
marine food webs
ca. 5 links
land vs ocean photosynthesis products
L: high light high nutrient low CO2 low water O: opposite
Diatom size
2 um - 4mm
diatom characteristics
10000+ spp.
abundant at high lats
single or chains
diatom classification
centric
pennate
diatom body form
hypotheca inside epitheca (frustules)
chloro., nucleus, oil
oil, projections, perforations
increased SA - buoyancy
Dinoflagellate size
2 um - 2 mm
dinoflag. characteristics
1000+ spp. usually solitary primitive plastids + secondary pigmants asexual -starch and lipids -mixotrophic -dont need Si -low SA:V -can migrate vertically
dinoflag. form
- armored or unarmored
- 1+ layers of cellulose
- 2 flagella in grooves
dinoflag. flagella grooves
cingulum- encircles, for rotation
sulcus - displacement
dinoflag. pigments
chlorophyll a, c
beta-carotene
peridinin
dinoflag. vs diatoms
- advantage over diatoms
- more abundant in tropic water
specialized dinoflag.
zooxanthellae
HABs
zooxanthellae
no flagella
symbiont in many species (coral, jellies, molluscs)
HABs
harmful algal blooms
- produce toxins
- deplete oxygen
- paralytic shellfish poisoning
types of Haptophytes
coccolithophores (most)
haptophyceae
haptophyte characteristics
370 spp 2-20um 1-2 chromatophore 2 flag. calcareous plates auto, hereo, mixotrophic
haptophytes responsible for
40% of carbonate production in modern seas
haptophyte speacilized structure
haptonema
defense or prey capture
sticky tip
Cyanobacteria character
prokaryote
blue-green algae
single, colony, filaments
starch, lipids
cyanobacteria well adapted to
nutrient-poor open ocean tropics
-like lots of sun and O2
cyanobacteria pigments
chlorophyll a, b beta-carotene xanthophylls phycoerytherin phycocyanin
carotenoids
beta-carotene (yellow)
xanthophylls (brown)
phycobilins
phycoerythrin (red)
phycocyanin (blue)
Nitrogen fixation
conversion of atm N into useable form
Nitrification
conversion of ammonia from waste and detritus to nitrate ions
Heterocyst
- contain specialized enzymes
- cyanobacteria
- N fixation / nitrification
Foraminiferan
zooplankton pseudopods multi-chamber test consume diatoms, bacteria sexual and asexual
foramin pseuodopods
form reticulopods - net-like structures
foramin limitations
2000 m
CCD
radiolarian
Actinopoda
- benthic grazer or planktonic suspension
- long needle-like pseudopods
- Si skeleton and spines
copepod
<1mm - few mm
- jerky motions
- large antennae
- complex life cycle
copepod feeding
create water stream w/ head appendages - moves particle down ventral surface - capture with 2nd maxillae - brought to mouth
copepod life cycle
6X Nauplii
5X copepodid
Major zooplankton
krill cladoceran foraminiferan radiolarian ctenophore arrow worm scyphozoan (jelly) siphonophore (MoW)
amount of E lost at each transfer
80-95%
C =
energy ingested
= A + F
= E assimilated + E lost Feces
A =
E assimilated
= P + R + U
= 2dary product. (growth) + E loss respiration + E loss nitrog. waste
P =
growth =
C - R - U - F
= food - respiration - urea - feces
force required to separate water molecules allowing organism to pass
viscosity
viscosity, T
negatively correlated
viscosity, S
positively correlated
SR =
sinking rate = (W1 - W2) / R V W1 = org density W2 = SW density R = surface of resistance V = viscosity of SW
why phytopl. should float or sink
nutrients
sunlight (good and bad if too high)
dont get stuck under thermocline
why zoopl. should float or sink
follow the phyto
flotation mechanisms
weight reduction
∆ surface of resistance
exploit water movement
reduction of weight, flotation
alter body fluid comp. (ammonium chl)
gas-filled floats
use lighter fluids (lipids)
changes in surface resistance, flotation
small (higher SA:V)
flattened shape
spines, projections
why smaller size more important for tropical plankton
higher T = less dense water
also more serious thermocline to watch out for
exploitation of water movement, flotation
langmuir convection
how langmuir works
day - heat, night - cool, T-driven convection cells
local (m’s - 100s m’s)
wind > 3 m/s
langmuir upwelling
convergent cells
tropic plankton pattern
low and relatively even all year
very little lag or difference
polar plankton pattern
one peak for each in summer
lag between phyto., zoop
Intensity of light at depth, I_z
= I_0 e ^ -kz
I_0 = light at surface
k = extinction coeffic.
z = depth
light attenuation curve
exponential decrease with depth
depth where
respiration = photosynthesis
compensation depth
no growth
(based on light)
phytopl. must remain above
temperate plankton pattern
two phyto. peaks (spring, fall)
spring peak bigger
zoop lag
depth of light penetration
absorption wavelength reflection scattering latitude season
peak wavelength penetration in water
blue
compensation depth depends on
latitude season sea surface conditions water clarity type of PP position relative to shore
strong thermocline all year
tropics
nekton selection
- mobility
- nervous, sensory systems
- fast swimming
- camouflage
- floatation
no connection or duct between the swim-bladder and the intestinal tract
physoclistous
physoclistous air control
specialised structures called the gas gland and ovale respectively
rete mirabile means
‘wonderful net’
latin
Fish with a connection (pneumatic duct) between the gas bladder and the esophagus
physostomus
rete mirabile is
counter-current exchange
cappillaries
allow gas uptake in fish with swim bladder
causes diffusion in rete mirabile
oxygen tension greater in venous than arterial blood
physostomus air control
via the mouth
nekton adaptations for buoyancy
- swim bladder
- swim fast
- gas filled cavity
- lipids
nekton, swimming fast
avoid sinking
streamlined
strong tail
EX. nekton quick swimmer
bonito
mackerel
nekton with gas-filled cavities
mammals (seals)
birds
nekton with lipids
fast fish (lipid-filled liver of shark) mammals (blubber)
nekton adaptations to surface of resistance
streamlined
long, thin
resistance to movement
frictional resistance
form resistance
induced drag
minimal frictional resistance
in spherical objects
minimal form resistance
in long thin object
proportional to cross-sectional area
induced drag increases
with speed or size
why does induced drag increase
laminar flow disrupted
forms vortices, eddies
Aspect ratio =
Height of caudal fin^2 / Area of caudal fin
fastest fishes caudal fin
high AR
therefore tall but narrow
Nekton adaptations, defense
Ventral keel
cryptic coloration
ventral keel
sharp angeled ventral edge allows light to illuminate ventral side and reduce shadow and visibility
nekton adaptations, sensory systems
lateral lines
ampullae of lorenzini
vision, hearing, olfaction
lateral line
canals length of fish body and over head
-detect pressure (movement)
ampullae of lorenzini
organ that can detect electrical signals in water (sharks, cartilagenous fish)
nekton need adaptations for body heat because
water has a higher thermal conductivity than air
nekton adaptation, heat
large (SA:V)
fat (blubber)
modified circulatory system
nekton modified circulatory system
warm arterial blood transfers heat to cooler venous blood; recycles heat; keep heat in organism core