Midterm 1 Flashcards

1
Q

Vegetation patterns exist in the tundra. Describe: Tussock tundra

A

Tussock tundra – special type of sedge meadow derived from dense clumps of the cottongrass Eriophorum vaginatum
• Comprise a large portion of carbon stocks

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2
Q

There are 3 zones of the boreal forest, list them.

A
  1. Closed boreal forest (southern)
  2. Open boreal forest (northern)
  3. Forest-tundra ecotone
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3
Q

In terms of sea ice, define: Shore-fast ice.

A

coastal regions where ice grows out from and stays attached to the shore

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4
Q

Describe this type of tundra tree: Krummholz

A

Krummholz – stunted wind-blown trees due continual exposure to fierce, freezing winds causes vegetation to become deformed

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5
Q

Conifers dominate the taiga for many reasons, describe how being an accumulator species contributes.

A

Most conifers are accumulator species – hoard minerals
• accumulated nutrients aren’t recycled – don’t lose leaves
• fires are often needed to break down dead plant material

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6
Q

Vegetation patterns exist in the tundra. Describe: Sedge meadows.

A

Sedge meadows: flat or rolling terrain dominated by a mat of grass-like sedge plants
• dense green swards or lawns in imperfectly drained lowlands, dominated by a variety of sedges (Carex sp) and cottongrasses (Eriophorum sp)
•Wettest after snowmelt – drier as summer goes on •Degrade with water pollution and grazing
• related to the open peatlands of the northern boreal forest

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7
Q

What 4 things (basic) are needed for snow crystals to form?

A

Water Vapor + Ice Nuclei + Cloud Droplets + Temperature <4°C

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8
Q

Define: Kettle hole.

A

A hollow created when buried blocks of glacier ice melt out.

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9
Q

Conifers dominate the taiga for many reasons, describe how mutalism with mycorrhizae. contributes.

A

Most taiga plants have mycorrhizae – interweavings of fungal mycelium and root tissue.

Mycorrhizae are a mutualism between plants and fungi.
• the plant provides the fungus with carbohydrates
• the fungus helps the plant get nutrients from the soil, like nitrogen

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10
Q

Describe the Pleistocene Epoch in terms of:

  • How many glacial retreats/advances?
  • How thick was the ice and how much land was covered?
A

A glacial period that had up to 20 glacial advances and retreats. There were massive ice sheets (3-4km thick) that covered 1/3 of earth’s land masses.

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11
Q

List the 5 major ice ages that earth has experienced (in order).

A
  1. Huronian.
  2. Cryogenian.
  3. Andean-Saharn.
  4. Karoo.
  5. Quatenary.
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12
Q
  • Reading by Dredge*
  • What kind of ground was present in the natural environment of Northern Manitoba (pre-ice age). How was this ground formed?
A

Northern Manitoba (Canadian shield) is a mix of ancient remnants of the earth’s original crust and newer rocks. Old crust was folded through mountain building cycles, while newer crusts was created by volcanic activity (sediments). Most of Churchill’s topography developed in the Late Precambrian time. Limestone makes up much of the sedimentary rock, and is made up of silt or clay sediments and skeletons of calcareous marine organisms.

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13
Q

What are ice wedges, and how does permafrost make them form?

A
Cold ground contracts in winter, cracks
• Cracks fill with water in spring
• Freezes and expanded by surrounding permafrost
• Forms ice wedges
• Repeat. Wedges grow over
time.
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14
Q

Are bogs and muskeg’s low or high in nutrients? What is their environment like (temp and nutrients)?

A
  • Low nutrients, but some plants are adapted to this and the chilly soils.
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15
Q

Describe this principle force acting on sea ice:

  • Ocean Currents
A

typically act in the opposite direction of the wind force
• act as a drag on the wind-driven sea ice motion
• important in longer-term ice motion (monthly to
yearly)

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16
Q

Eastern and northern Manitoba’s Boreal Forest and peatlands are exceptionally rich in ___.

A

Eastern and northern Manitoba’s Boreal Forest and peatlands are exceptionally rich in carbon

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17
Q

Define: Albedo.

What does this influence?

A

% solar energy reflected back into space (big influence on surface energy).

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18
Q

How much of the world’s freshwater do cryospheres hold?

A

75%

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19
Q

Describe ice nucleation active (INA) bacteria.

A

Presence on plant surfaces causes ice nucleation at temperatures 5-8° warmer than in the absence of the INA bacteria.

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20
Q
  • Reading by Dredge*

- Today, what is the climate of Churchill (basic)? Is it colder or warmer than previous times, and how can we know this?

A
  • Marine subarctic climate. Most of northeast Manitoba has continuous permafrost (up to 80m thick in Churchill).
  • There is evidence of tree migration when warm climates prevailed, and it is believed that trees were present 300km more north compared to now.
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21
Q

Optimal function of enzymes is __-__ C.

A

Optimal function of enzymes is 37-40 C.

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22
Q

Define: Tree line.

A

Treeline: northern boundary of the forest-tundra zone
• the zone where ‘forest with patches of tundra’ merges
into ‘tundra with patches of forest’
• the northern limit of the growth of trees that >5m high
• No distinct boundary - gradual change

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23
Q

Has carbon energy pathway to benthic fauna changed over time? Explain this by answering the following question using the Atlantic walrus (a bioindicator).

These walruses are..
• Primarily benthivores
• Can consume 57kg of bivalves per day
• Central place foragers
• Haul-out in large aggregations

2) Is there latitudinal variation associated with different sea ice phenology?

(pretty much just yes or no)

A

Dependent on latitude and sea ice conditions • Jones Sound – no change, other areas had change.

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24
Q

Boreal forest areas burning is highly episodic with high inter annual variability. Approximately 0.2 - 7.6 million ha are burnt a year. Is this increasing over time?

A

Yes, and will likely continue to increase with more climate change.
- There are more HUGE fires now compared to before (more area burnt), but smaller fires haven’t changed as much.

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25
Q

First year ice vs multi year ice.

A
  • First year ice forms in grades every year.

- Multi-year ice is a permanent ice zone, may contain younger ice and open water

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26
Q

There are 3 types of fires, surface, crown, and ground fires.

Describe surface fires in terms of what burns/where.

A

Surface fires: burn fuels at the ground surface

• shrubs, grasses, fallen branches, litter

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27
Q

How do invasive species post a threat to forests?

A

Invasive species can pose an even greater threat
• no predators, diseases or other natural forces in
Canada to control their population

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28
Q

Define: Minerotrophic.

A

Fens are minerotrophic – water supply comes mainly from streams or springs.

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29
Q

How are peatlands are growing points for permafrost throughout the taiga?

A

In summer, sphagnum dries out, insulates against incoming heat
•Sphagnum soils are among the last to thaw in the boreal forest.
In autumn, heavy rains and freeze-thaw cycles: sphagnum becomes water-saturated – conducts outgoing heat

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30
Q

List 5 principle forces acting on sea ice.

A
  1. Wind
  2. Ocean Currents
  3. Coriolis Force
  4. Internal Ice Stress
  5. Sea Surface Tilt
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31
Q

CO2 levels are increasing by __% per year (anthropogenic cause).

A

1.5%

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32
Q

How does permafrost freeze in winter (ie. from top or bottom?), and how does it thaw? What happens after it thaws?

A
  • Active layer freezes in winter, thaws in summer (top-down)
  • After active layer thaws, the water can’t drain
  • Can have very wet and boggy ground in summer
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33
Q
  • practice exam question *

T or F - Is the rate of ocean heat content now increasing faster?

A

True!

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34
Q

How (and when) was Lake Agassiz formed?

A
  • Ice sheets blocked meltwater from moving out into Hudson’s bay, and formed a freshwater lake.
  • The lake covered most of the mid continent (including Manitoba) for ~5000 years.
  • Emptied 8000 years ago when the last ice sheet disintegrated, but left areas depressed by water and ice.
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35
Q

One theory for the reason of more autochthonous lake input (“stuff” coming in from farther places) may be due to more run-off. Is there evidence for increased precipitation, and how would this cause the problem?

A

• Significant increases in total annual precipitation detected; Trend driven by summer rainfall.
- Precipitation is increasing by +17-19 mm per decade.

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36
Q

List the 3 layers of the nivean environment.

A
  1. Subnivean
  2. Intranivean
  3. Supranivean
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37
Q

What do climate change models predict in terms of insect outbreaks? What would be detrimental to insect outbreaks (ie. prevent them)?

A
  • Increased temperature increases could increase the risk of future beetle outbreaks
  • esp. at high altitudes and latitudes where not as common currently

• Extreme winter weather patterns (e.g., large temperature drops) are detrimental to beetle outbreaks

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38
Q

Describe viruses in sea ice (part of food web).

A
  • High ratio of viruses to bacterias, likely indicating high infection rates but limited information;.
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39
Q

Describe this area within the boreal zone in terms of trees (basic): Forest-tundra ecotone.

A

transition between

taiga and tundra

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40
Q

What kind of camera can be used to detect infrared (like an animal)?

A
  • FLIR infrared camera.
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41
Q

In terms of glaciers, what is plucking?

A

Glacier picks up rocks and sediment and deposits it elsewhere.

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42
Q

What forms corn snow?

A
  • meltwater or rain enters the snowpack

- Thawing and freezing cycle = increased vapor migration and freezing lead to large rounded grains

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43
Q

Snow depth/ melt pond coverage largely control light availability in sea ice.

How does attenuation doffer between snow and ice? What is attenuation?

A
  • Snow – 15 m^-1
  • Ice – 2m^-1

Attenuation is how easily light can penetrate something. Snow increases attenuation (less light scatter, less light for algal growth).

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44
Q
  • Reading by Dredge*

- Describe how Churchill evolved with its 3 major glaciations.

A

1st: The ice melting created a large lake/sea with clay deposits. Eventually land rose and water receded, leaving tundra-vegetation to cover the area.
2nd: Ice again formed and melted, leaving a sea and vegetation similar to what is seen today (tundra moss, shrub-sedge, birch communities) and animals like woolly mammoths, beavers, snails, and molluscs occupied the area.
3rd: This glaciation brought fossiliferous limestone debris and created the Glacial Lake Agassiz which covered most of the mid-continent.

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45
Q

In terms of formation of snow pack, define: Constructive metamorphism (temperature gradient metamorphism).

A

New crystals are formed
• Water vapor rises – moves from warmer area near ground
• Water vapor crystalizes as it hits colder temperatures
• With a greater temperature gradient, more vapor is transferred
• More tightly packed near top • Bottom snow layers becomes
loose, less dense, creating crystalline snow = depth hoar

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46
Q

There are 3 types of fires, surface, crown, and ground fires.

Describe ground fires in terms of what burns, and how they form.

A

Ground fires: burn subsurface organic fuels by smoldering combustion
• often ignited by surface fires
• found in deep peat deposits in a peat bog
• burns most intensely & spreads fastest at the lowest levels
• can move very fast
• may smolder for days to months….then break out on the surface to be fanned by winds into full-scale crown fires

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47
Q

Impacts of disturbances are influenced by spatial factors, temporal, and magnitude.

Describe magnitude.

A

• Intensity - Physical force of the event

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48
Q

Describe this type of tundra tree: Flagged trees

A

Flagged trees – branches grow on only one side, due to abrasion by wind and blowing ice crystals.

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49
Q

Reading by Pruitt

Describe this quality of snow cover that affects living organisms: Density.

How does it affect animals (3)?

A
  1. It expresses the water content of the snow cover, so dense snow will cause more spring run off.
  2. Higher density can mean higher insulation for subnivian organisms.
  3. The insulating properties of density makes the pukak layer warmer than the upper layers, allowing animal and bacterial activity to proceed (which increases carbon dioxide up to 5x more than outside).
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50
Q

To stay alive in cold, life has a few options: Stay above 0C, or drop below 0C.

In terms of staying above 0C, describe staying warm for endotherms.

A
  • Adjust body temperature.
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51
Q

What is the microbial loop in sea ice primary production? How important are bacteria?

A
  • ≈50% of sea ice primary production is in the form of dissolved organic matter (DOM).
  • DOM is the main substrate for bacterial growth.
  • Microbial loop: Bacteria assimilate DOM into the biogenic particulate fraction that can be utilized by bacterivores, transferring carbon back into the classical food web.
  • Bacterial respiration has been estimated at 11% of primary production.
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52
Q

Define: Peat plateaus.

Size?

A

Flat-topped areas of peat (smaller).

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53
Q

What is a consequence of younger and thinner ice (old ice is melting)?

A
  • more light transmitted through the ice = positive feedback loop of more warming
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54
Q

The Anaktuvuk river fire was the largest fire on record for the tundra biome (2007), where the tundra ecosystem lost >2 kg C/m^2. What is the problem with so much carbon being released?

A

Overall, the fire released ~2.1 metric tons of C to the atmosphere

• an amount similar to the annual net C sink for the entire Arctic
tundra biome

Carbon balance will subsequently be altered due to
• reduced albedo (absorbs more energy)
• removal of soil organic matter (less insulation)
= Thaws permafrost

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55
Q

What are pingos, and how does permafrost make them form? What are they similar to?

A

Subsurface water can freeze and force frozen ground upward (pressure) to form cone-shaped mounds with cores of ice.
Similar to palsas, but are much bigger and they grow below the active layer (vs in the active layer).

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56
Q

What is the main cause of ice ages (very brief)?

A

Changes in thermal energy.

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57
Q

In ice as a biome, who are the algal primary producers and how much of the biomass are they? Does it change from first bloom to melt and late season?

A
  • Pennate diatoms dominate bottom ice algal bloom biomass in the Arctic, can make up >90% biomass in the bottom ice algae community of landfast ice.
  • During melt, autotrophic flagellates tend to dominate. Sub-ice communities can also develop in late season (e.g., dominated by the chain-forming centric diatom Melosira arctica
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58
Q

How do warmer summer temperatures affect fish who like cold water (trouts, walleye, Cisco)?

A

Species are using deeper, colder water in Ontario lake.

Nearshore (littoral coupling) is lower with higher summer temperatures.

Found in deeper waters when littoral zone warmer.

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59
Q

Moose were found to not avoid roads/human activity. They did however, avoid deer. Why?

A

Brain worm – carried by deer in brain
• Fatal in moose

Larvae passed through blood into feces – then onto snails or slugs Causes sickness in white-tailed deer

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60
Q

Sea ice is one of the largest biomes, and it’s extent equals __-__% of the world’s ocean surface at any one point in time.

A

Sea ice extent equals 4-6% of the world’s ocean surface at any one point in time.

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61
Q

List 2 ways/methods (general, not subcategories) boreal plants escape frost damage.

A
  1. Extracellular (aka intercellular) freezing, which prevents intracellular freezing.
  2. Supercooling.
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62
Q

Describe this quality of snow cover that affects living organisms: Hardness.

What influences it?
Give an example of an animals it effects?

A

primarily influenced by wind and thaw; force required to collapse the api.

Affects terrestrial mammals like caribou.

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63
Q
  • Reading by Warner *

Canadian peatlands cover about __% of the land surface

A

Canadian peatlands cover about 13% of the land surface.

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64
Q
  • practice exam question *

New data suggests that the ocean heat content is now rising much faster than before. By how much? (%).

A

50-75%

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65
Q

Canada has the most surface water of any other country, what are these surface waters critically important to?

A

• All life and the maintenance of biodiversity on the
landscape
• A large range of ecosystem goods and services
(e.g. drinking water, hydro power, agriculture, sanitation, commercial fisheries, recreation), big economic effect too

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66
Q

List the 3 most abundant greenhouse gases, and how long each stays in the atmosphere.

A
  1. Water vapour (9 days)
  2. CO2 (up to 100 years)
  3. Methane (12 years)
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67
Q

Boreal areas can also be adapted to disturbance, what do larger scale disturbances create, and what animals can live here?

A

Larger disturbances, over time, create habitat for species that prefer large, uniform forests
• marten, woodland caribou, & great grey owl

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68
Q

Do bigger or smaller animals have a higher mass specific metabolism?

A

smaller animals.

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69
Q

Why do conifers dominate the taiga? List their 6 main adaptions.

A
  1. Short growing season.
  2. Conical shape, dark colour.
  3. Low surface area.
  4. Tolerate nutrient poor soils.
  5. Accumulator species.
  6. Mutalism with mycorrhizae.
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70
Q

Describe this principle force acting on sea ice:

  • Coriolis Force
A

causes objects to accelerate because of the rotation of the earth.
• affect processes that occur at the global scale, such as
ocean currents, winds, and ice motion
• in the Northern Hemisphere, causes objects to deflect
to the right
• increases toward the poles – plays an important role in determining sea ice motion

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71
Q

How are fens “fed”? Describe this. What “types” can they be?

A

Fens are minerotrophic – water supply comes mainly from streams or springs.
Fens can be peatlands & mineral wetlands.

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72
Q

Damage can be caused by _____ which feed & reproduce in the inner bark of trees; causes tree mortality. They essentially girdle the tree.

A

bark beetles (Scolytidae)

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73
Q

Radiant energy from the sun warms the earth’s atmosphere and surface (water and soil) and becomes heat energy. Describe this mechanism by which energy is transferred through the atmosphere:

  • Conduction
A
  • Molecule to molecule (or solid to solid) exchange.
  • Depends on the temperature gradient, and rate of transfer varies by substance.
  • Air is a poor thermal conductor, while snow is a poor conductor.
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74
Q

When did Canada develop peatlands? Do they span tot he high arctic?

A

Canada’s extensive peatlands developed since the last glaciation
< 10,000 years ago.
Peatlands are rare in the High Arctic because climatic aridity and harsh growing conditions are not particularly conducive to peatland formation

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75
Q

How does wind speed influence damage? Are damaged trees more susceptible to other problems?

A

Wind speed largely determines the extent of damage to trees
• Larger trees are more susceptible to windthrow - lever
• stands that have been recently burned

Damage to tree stands increases the probability of pathogen outbreak and tree mortality

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76
Q

Radiant energy from the sun warms the earth’s atmosphere and surface (water and soil) and becomes heat energy. Describe this mechanism by which energy is transferred through the atmosphere:

  • Radiation
A
  • Transfer of energy by electromagnetic waves (ie. sun in form of visible light).
  • Wavelength is related to vibration frequency: Long - infrared, short - ultraviolet.
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77
Q
  • Reading by Warwick *

In terms of the ecological implications of changes in the arctic cryosphere, what effect does the warming arctic cryosphere have on: Permafrost.

  • Effect of more water, effect on vegetation and life once it dries up, contaminants.
A
  • Erosion will release a lot of water that will ruin forests and turn them into fens, and can create many lakes and wetlands.
  • Southern invasive species.
  • Increased vegetation biomass with drier conditions can increase fires.
  • Thawing permafrost releases organic matter, gasses, inorganic solutes, and microbiota into the environment.
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78
Q

What is the high arctic in terms of:

  • General area (relative to tundra)
  • Vegetation, ground, amount of snow
  • Winter thaw or no?
A
  • northern tundra
  • vegetation is short
  • A lot of bare ground
  • less snow
  • No winter thaw
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79
Q

How do amphipods combat freezing?

A

To combat freezing ice amphipods conform osmotically to their environment

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80
Q

Animals in the arctic follow the SA/vol rule called the: __/__ power law

A

3⁄4 power law

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81
Q

Reading by Carlson + Windsor

  • How long ago did the last ice age occur?
  • How does the melting of glaciers differ for glaciers on land (coastal vs inner) and on the ocean?
A
  • 2.6 mya, ice sheets covered large portions of the northern hemisphere, which melted rapidly in the southernmost land margins, and retreated slower on land. Marine ice sheets had a delayed onset of retreat relative to temperature (retreat followed by rapid collapse).
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82
Q

Define: Biome.

A
  • A major geographical area characterized by a particular type of flora and fauna – primarily determined by CLIMATE.
  • A large-scale community of organisms shaped by common environmental conditions (climate and geology)
    • usually named after its predominant vegetation association
    • general and global, encompassing numerous smaller- scale ecosystems
    • e.g., Amazon Basin rainforest vs Congo Basin rainforest
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83
Q

How long ago did this ice age occur?

- 1/5: Huronian

A

2.4 to 2.1 billion years ago.

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84
Q

Describe this Milankovitch cycle explaining changes in insolation reaching earth:
- Tilt.

  • How does it vary, and how long is the cycle?
A
  • Axial obliquity: Ranges from degrees of 22.1 to 24.5 (currently 23.4 and decreasing).
  • 41k year cycle.
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85
Q

Define: Supercooling.

A

lowering the temperature of a liquid or a gas below its potential freezing point, without it becoming a solid
= Ice crystals cannot form at 0°C

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86
Q

What is subsea permafrost, and how did it form?

A

Under the Arctic ocean, not in Southern hemisphere Formed >11,000 years ago, during the last ice age
• Lower sea level
• Exposed land froze
When ice sheets melted, sea level rose & covered up areas of permafrost
Today, some of seafloor still frozen up to 100m thick under seafloor

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87
Q

Milankovitch cycles explain changes in insolation reaching earth. List the 3 different cycles/changes in orbit.

A
  1. Eccentricity.
  2. Tilt.
  3. Precession.
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88
Q

Where do peatlands occur and how much of this region do they cover globally?

A

Peatlands occur mainly in boreal regions, covering 25-30% of the boreal forest region globally

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89
Q

How does the cryospheres influence the world?

A

Influences global energy balance and ecological functions of northern environments

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90
Q

In terms of formation of snow pack, define: Destructive metamorphism (equi-temperature).

A

“destruction” of individual grains as snow settles (ie. by wind)
• Decrease in volume
• Increase in density (reduced space between
crystals)
• Occurs early in snow settling

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91
Q

Define: Talik.

A

Soil or unfrozen ground, year-round, can be above, within or below permafrost

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92
Q

___ can be a major fire season.

A

Spring.

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93
Q

Boreal forest areas burning is highly episodic with high inter annual variability. Approximately how many million hectares are burnt a year (the range)?

A

0.2 - 7.6 million ha.

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94
Q

Why do carbon concentrations in the atmosphere oscillate?

A

They go down a bit in summer due to increased plant photosynthesis.

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95
Q

Main grazers of ice algae include heterotrophic protists (microfauna), metazoans that live within the sea ice (meiofauna), and under-ice metazoans.

Describe the meiofauna of ice. What are the main ones, and how much algal life (and daily ice algae production) do they consume?

A
  • Dominated by flatworms, crustaceans (including harpacticoid and cyclopoid copepods), nematodes, rotifers, and bentho- sympagic larvae
  • Consume about 1% of the ice algae standing stock and <6% of daily ice algae production
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96
Q

What is the most widespread peatland in Canada?

a. fen
b. bog
c. muskeg
d. swamp
e. tundra

A

B - bog

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97
Q

Are we in a new epoch, and headed toward a “hothouse earth”?

A
  • It is possible that we are driving the glacial-interglacial limit cycle toward a hotter state…
  • Even if the earth temperature stabilized, it’s still hotter than at any time modern humans have existed.
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98
Q

How long ago did this ice age occur?

- 2/5: Cryogenian

A

720 - 630 million years ago.

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99
Q

Lightening fires are destructive in the boreal forest. They make up:

–__% of total fires
–represent __% of area burned

A

Lightening fires are destructive in the boreal forest. They make up:

–35% of total fires
–represent 85% of area burned

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100
Q

Impacts of disturbances are influenced by spatial factors, temporal, and magnitude.

Describe temporal factors (3).

A
  • Frequency - number of events per unit of time
  • Duration
  • Time of year
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101
Q

Describe how slopes can influence the amount of frozen ground present.

A
  • slope facing towards sun get more sunlight (radiation) – facing south in northern regions
  • warmer ground = less likely to freeze (angles of insolation)
  • steep slopes do not get as much direct sunlight = relatively colder, also don’t hold as much snow
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102
Q

Animals may experience severe range contractions and possible extinction due to environmental change… what 2 things can occur due to this? (Ie. genetically).

A
  • Isolation.

- Genetic drift.

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103
Q

In North America, what kind of forests would you find north?

A

Tundra.

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104
Q

Describe this principle force acting on sea ice:

  • Wind

What does it cause/how?

A
primary force responsible for ice motion
• Short-term ice motion – days to weeks
• causes the ice to drift – drag force
• force depends on the speed of the wind and the
characteristics of the sea ice surface
• rough ice surface is affected more
• Speed is predictable - sea ice that drifts freely
moves at 2% of the wind speed
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105
Q

Where would a jack pine be found (habitat)? Is it common in MB?

A
Jack pine (Pinus banksiana): continental, found on dry, sandy outwash plains and former dune areas – low nutrient, droughty substrates not tolerated by spruce and fir.
• most common pine in Manitoba
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106
Q

Do boreal ecosystems experience natural disturbances often? What are 4 major ones?

A

Boreal ecosystems are regularly subjected to
large-scale natural disturbances - they’re disturbance-based ecosystems:

Major disturbances:

1) Fire
2) Insect infestations
3) Disease
4) Severe weather

Boreal forest ecosystems are constantly changing Most boreal vegetation and wildlife species have adapted to, and depend on, disturbance.

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107
Q

Define: Discontinuous permafrost.

How thick is it?

A

Discontinuous→ occurs under 50-90% of land in a defined area

• Up to 10 m thick

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108
Q

What is the intermediate disturbance hypothesis?

A
  • Highest diversity is often maintained at an intermediate level of disturbance.
  • Both frequency and scale of disturbance affect diversity
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109
Q

How does dried peat have some uses for humans?

A

Dried peat is a traditional fuel in places where it is of high quality and easily accessible (e.g., Ireland).
- Also in alchohol!

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110
Q

Fire size in the boreal forest varies:

• Most fires are small (___% are ___ ha
these together represent 97% of area burned

A

Fire size in the boreal forest varies:

• Most fires are small (70% are <5 ha)
• only 3% of fires are >200 ha
these together represent 97% of area burned

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111
Q

__% of land (globally) has permafrost underneath. Over __% of Canada does.

A

24% of land (globally) has permafrost underneath. Over 50% of Canada does.

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112
Q

Why is it colder at the poles of the earth (3 main reasons)?

A
  • When sunlight hits the ground obliquely (at an angle), there’s more surface area to cover and less absorption, so less amount of energy per unit area.
  • The sun’s rays have to travel farther through the atmosphere at higher latitudes, resulting in less heat reaching the ground, less absorbed into the ground, and more reflected into the atmosphere.
  • Long nights in winter (months sometimes), more heat is lost to the atmosphere than gained at the surface.
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113
Q

What 3 things can animals do if the environmental conditions change outside their tolerance?

A
  1. Move (new location).
  2. Adapt (adapt to new conditions in original habitat).
  3. Go extinct.
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114
Q

T or F: fires can cause a big decrease in forest soil depth.

A

True

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115
Q

Why can’t trees grow in the far north?

A

The lack of summer warmth – not cold winters.
• Photochemical reactions (photosynthesis) can
only happen if plant tissues are warm enough
• Results in low growth (leaves and roots are a necessity, wood/growth is a luxury).

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116
Q

What is Clements’ idea of a “superorganism”?

A
  • Communities are “superorganisms”)
  • Demonstrates that communities change over time in very discrete ways (like human development) ultimately resulting in a predictable endpoint
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117
Q

Does any flora or fauna live on glaciers and ice sheets?

A
  • Microbes can live on the ice surface or in solid ice.
    Microbe – any living organism that spends its life at a size too tiny to be seen with the naked eye.
    ‒ includes bacteria and archaebacteria, protists, some fungi and even some very tiny animals
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118
Q

What’s the difference between a homeotherm and endotherm?

A

Homeotherm: animal that maintains an approximately constant body temperature.

Endotherm: generates heat to maintain its body temperature. They rely primarily on their own energy reserves to produce heat in cold conditions.

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119
Q

List a few things that can influence albedos.

A

Snow depth, age, vegetation, sunlight, cloud cover. Pollution (adds more cloud nuclei and changes reflection/absorption - thicker = more absorptive).

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120
Q

Main grazers of ice algae include heterotrophic protists (microfauna), metazoans that live within the sea ice (meiofauna), and under-ice metazoans.

Describe the under ice metazoans. What are the main ones, and how much algal life (and daily ice algae production) do they consume?

A
  • Dominated by gammarid ice amphipods and planktonic calanoid copepods
  • Amphipod grazing rates estimated at 1-3% of ice algae
  • Calanoid copepods observed to graze on ice bottom, but no direct estimates to date
  • > 65% sloughed ice algae floats or sinks slowly through the water column where it is primarily grazed by copepods
  • Gelatinous zooplankton are commonly observed feeding along the sub-ice environment
  • Arctic cod is a keystone species, providing a central link between grazers and higher trophic level fish, birds, and marine mammals
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121
Q

What are 7 things that can influence the amount of frozen ground?

A
  • Soil type
  • Lakes and rivers
  • Snow
  • Slopes
  • Wind direction
  • Plants
  • Peat
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122
Q

How can plants accomplish extracellular freezing?

A
  • “Antifreeze” accumulates inside cell walls (sap and other compounds)
  • Ice crystals form in between cells in intercellular spaces
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123
Q

Is there a correlation between amount of peatland on a landscape and fire events? What else correlates to more fire events (relating to peatland)?

A
  • Yes!

- Drought.

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124
Q
  • Reading by Warwick *

In terms of the ecological implications of changes in the arctic cryosphere, what effect does the warming arctic cryosphere have on: Lake and river ice.

  • Increases in certain organisms, contaminants, invasive species.
A
  • Less wind-induced mixing + more light for photosynthesis, will cause increases in zooplankton and contaminant transfer to high trophic levels.
  • Shifts habitat of cold adapted species.
  • Increases in mercury and other contaminants in fish.
  • Southern plants and fish can invade.
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125
Q

How does the albedo in lake vs sea ice differ? What does this have to do with their type of growth?

A
  • Albedo is much higher in sea ice due to the numerous impurities found in sea ice (lamellar growth) compared to lake ice (planar growth).
  • 99.9% of impurities are expelled from accreting lake ice
  • 10 to 40% impurities are trapped within sea ice
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126
Q

What type of forest fires usually born most intensely and spread fastest?

A

Ground fires.

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127
Q

List 5 things that cause the formation of layers in snow.

A
  1. Wind→ increase wind, increase convection (primary driver)
  2. Water Vapor→ vapor gradient between snow and air
  3. Radiation→ Shortwave and longwave
  4. Advected Heat (Rain)
  5. Soil contact→ convection
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128
Q

Define: Climate.

A
  • Average state of Earth’s surface conditions.
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129
Q

In terms of sea ice, define: Pack ice.

A

almost constantly in motion, farther from shore, drifts in response to winds, currents, etc

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130
Q

What is a problem with arctic warming in terms of light reflection (albedo)?

A

The Arctic is warming twice as fast as the rest of the planet
• changing from a highly reflective surface to a very
dark surface
• Sunlight that would normally be reflected back into
space (by sea ice, snow or glaciers) is instead being absorbed by dark surfaces (ocean, land & rock)

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131
Q

What’s the most recent glaciation in North America?

A

Wisconsinan glaciation.

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132
Q

When Lake Agassiz was able to drain into Hudson’s bay, and the depressions left filled with marine water, this became the ______ ______.

A

Tyrrell Sea.

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133
Q

Fresh water is most dense at…

a. 0C
b. 2C
c. 4C
d. 6C
e. 8C

A

c - 4 degrees.

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134
Q

Describe how plants can influence the amount of frozen ground present.

A

• Keep soil cooler in summer – leaves can block
sunlight from reaching the ground
• especially evergreen trees
• create bare ground at their base – loses heat easily
• block sun in winter – more likely for frozen ground

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135
Q

One way to reduce heat loss is to get bigger. Describe Bergmann’s rule.

A

Animals of larger size are found in colder environments.

Less SA/vol ratio, less heat loss.

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136
Q

What happens to species that were around before the last glaciation?

A

Their range likely expanded and contracted over time.

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137
Q

Low temperatures cause rigidity of membrane phospholipid bi- layers resulting in a loss of ion permeability. Production of ______ fatty acids (e.g., ___-__) play a particularly important role to avoid this increase in viscosity.

A

Low temperatures cause rigidity of membrane phospholipid bi- layers resulting in a loss of ion permeability. Production of polyunsaturated fatty acids (e.g., omega-3) play a particularly important role to avoid this increase in viscosity.

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138
Q

What will happen to the tree line as the arctic warms?

A
  • Trees move north
  • Temperature-induced drought
  • Animals will follow!
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139
Q

How do insect outbreaks affect fire risk?

e.g. spruce budworm; Siberian silkworm

A
  • Early summer drought enhances the insect population.
  • Forest structure (suitable host trees) enhance insect population further.
  • Insect population reduces forest biomass
  • Forest structure (ex. dead trees) and biomass enhances fire hazard.
  • Fire hazards reduce forest structure and biomass.
  • Climate enhances fire hazard - it’s a cycle!
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140
Q

__% of the land in the Northern Hemisphere freezes seasonally.
• ~__ million square kilometers

A

58% of the land in the Northern Hemisphere freezes seasonally.
• ~55 million square kilometers

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141
Q

How can global warming change the carbon balance in the tundra?

A

Warming may change the carbon balance on the tundra – from a carbon sink to a carbon source

  • More carbon will be released due to decomposing microbes that release carbons, that won’t be offset by increased plant growth.
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142
Q

What is ecological succession?

A
  • Communities change through time.

* Succession is the replacement of one community by another.

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143
Q

Why are boreal forests so susceptible to large- scale insect disturbance? 2 main reasons.

A

1 Relatively lower species diversity = less predators

2. Relatively lower productivity

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144
Q

Describe Maritime snow in terms of:

  • General description
  • Snow depth
  • Layers of snow
A

Warm, wet, many melt features (ice etc..)

  • Snow depth: 75-500 cm
  • Layers of snow: >15
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145
Q

What kind of tree is a tamarack, and where would it be found?

A
  • Deciduous conifer that lives on thin, waterlogged substrate in level areas with permafrost.
  • Adapted to avoid water stress in spring.
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146
Q

What 2 things determine snow crystal shapes?

A
  1. Humidity

2. Temperature

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147
Q

There are 3 ways the sun’s energy can be reflected and absorbed, list the 3 (and relative proportion of each).

A
  1. Absorbed in the atmosphere (23%).
  2. Absorbed at the surface (48%).
  3. Reflected (29%).
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148
Q

Boreal forests are exceptionally unstable in vegetation. What’s most important cause of disturbance in boreal ecosystems?

A

Fire.

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149
Q

Define: Polynyas

A
  • Persistent (and predictable) open water surrounded by ice
  • sustained by winds or ocean heat
  • often occur near coasts, fast ice, or ice shelves
  • One of the major lifelines to Arctic ecosystem structure and function
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150
Q

How would an organism (ie. human) react to…

  • is below LCT
  • in TNZ
  • above UCZ
A
  • is below LCT: shivering
  • in TNZ: changes in blood flow (dnm?)
  • above UCZ: sweat
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151
Q

Are forests on the coasts more or less likely to burn? What other disturbances are also common here?

A

Forests on the coast are less likely to burn

But other impacts: wind & coastal erosion resulting in death of conifers

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152
Q

Are current CO2 concentrations very different from historic levels? Does it vary?

A

Over the past 20 million years, the Earth’s climate has oscillated between relatively warm and relatively cold conditions:
•During interglacial periods, atmospheric CO2 concentrations were relatively high
•during glacial periods, CO2 concentrations were relatively low.
We are currently in an interglacial warm period

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153
Q

To stay alive in cold, life has a few options: Stay above 0C, or drop below 0C.

In terms of staying above 0C, describe staying warm for ectotherms. Many possible options revolving around a common theme!

A

Ectotherms don’t produce their own heat, so must reduce heat loss (by habitat selection, move where it’s warmer).

  • Can move underground.
  • Communal nesting/huddling.
  • Alternative life stage (in many inverts, overwinter in egg stage, ie. mosquitos).
  • Move into deeper water.
  • Move under snow (subnivian layer)
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154
Q

Generally,
Ectotherm = _____
e.g., most insects, fish, herps

Endotherm = _____
e.g., most birds & mammals

A

Generally,
Ectotherm = Poikilotherm
e.g., most insects, fish, herps

Endotherm = Homeotherm e.g., most birds & mammals

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155
Q

What triggers insect population increases and associated widespread tree mortality? 2 reasons (causing 2 thing).

A
  • sustained periods of abnormally warm weather
  • drought

–> Both are correlated and create dry conditions. Multiple years of this can have a synergistic effect (much more damage than just 1 year).

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156
Q
  • practice exam question *

New data suggests that the ocean heat content is now rising much faster than before. By how much?

A. 10%
B. 50-75%
C. 100%
D. 150%
E. No change.
A

B

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157
Q

_____’s _____ forest is the largest remaining intact forest.

A

Canada’s boreal forest is the largest remaining intact forest.

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158
Q

Describe Tundra snow in terms of:

  • General description
  • Snow depth
  • Layers of snow
A

Thin, windblown, no trees, few melt effects

  • Snow depth: 10-75 cm
  • Layers of snow: 1-6
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159
Q

What was the largest fire on record for the tundra biome (2007)?

A

Anaktuvuk River fire in 2007
• burned > 1,000 km2
• largest fire on record for the tundra biome

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160
Q

Describe how a pitcher plant in a sphagnum bog can still collect nutrients.

A
  • Their red colour attracts flies and other meat-eating insects, they have hairs inside covered with a slippery gel.
  • The insects fall into the watery solution and are trapped and drown.
  • Plant secretes digestive enzymes into this warm, sun-heated solution; OR bacteria, microbes, and mosquito larva digest & the plant gets the waste products.
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161
Q

What is the definition of Boreal Ecology (very brief)?

A

Ecology of the north.

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162
Q

Describe this layer of a nivean environment, and give an example of an animal that may live here.

  • Intranivean
A

within the snow

• Area where some animals burrow, e.g. ruffed grouse

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163
Q

T or F: Canada is home to the world’s largest lakes and rivers.

A

True!

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164
Q

Main grazers of ice algae include heterotrophic protists (microfauna), metazoans that live within the sea ice (meiofauna), and under-ice metazoans.

Describe the microfauna of ice. What is the main one, and how much carbon is it consuming?

A
  • Consume ≈1% of total particulate carbon during an ice algae bloom and do not seem to consume diatoms (Michel et al. 2002)
  • Mainly bacterivores
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165
Q

Describe this quality of snow cover that affects living organisms: Density.

What influences it?

A

primarily influenced by wind and thaw; measured as weight divided by volume
• Expressed by water content

166
Q

Define: Tree line.

A

Treeline: northern boundary of the forest-tundra zone
• the zone where ‘forest with patches of tundra’ merges
into ‘tundra with patches of forest’
• the northern limit of the growth of trees that >5m high
• No distinct boundary - gradual

167
Q

Describe Alpine snow in terms of:

  • General description
  • Snow depth
  • Layers of snow
A

Intermediate to deep, depth hoar & wind crusts, some melt features

  • Snow depth: 75-120 cm
  • Layers of snow: >15
168
Q

Boreal areas can also be adapted to disturbance, and smaller scale disturbances may create an “__” habitat. What is this, and what animals can live here?

A

Smaller-scale disturbances create ‘edge’ habitat

• different forest types or landscape features come together
• habitat for species such as moose, black bear and ruffed
grouse

169
Q

What does the gyspy moth (invasive species/ alien) do? How were they introduced and why are they so detrimental?

A
  • Defoliates a wide range of tree species, particularly hardwoods.
  • Concentrations are notable in urban areas such as Saskatoon and Winnipeg
  • introduced to North America in the late 1860s by a French naturalist attempting to cross the European gypsy moth with North American silkworms. His intent was to establish a silk industry on this continent.
  • overwinters in egg masses attached to the bark of trees; may contain 100 - 1,000 eggs
170
Q

What 3 major changes occur in the environment as a consequence of glaciation?

A
  1. Change in climate patterns and zones (spatial and temporal).
  2. Change in location and extent of suitable habitat (varies by species).
  3. Routes of dispersal (Ice bridges vs water between land masses).
    - Overall, affects distribution of species.
171
Q

Define: Permafrost.

A

Frozen ground (@ or below 0°C) for > 2 yr.

172
Q

Define: The Suess effect in oceans.

A
  • Oceans are carbon sinks.
  • Most of world’s carbon is 12C, so environment is absorbing more of that isotope.
  • Changes in the ratio of atmospheric concentrations of the heavy and light isotopes of carbon are changing (due to increased CO2), there is less 13C and 14C now.
173
Q

____ is the bottom snow layer.

A

pukak

174
Q

How do algae counteract osmotic stress in ice?

A

To counteract the negative effects of osmotic stress on metabolism, ice algae accumulate organic osmolytes, also called “compatible solutes’’

175
Q

What is made when permafrost melts in the taiga?

A

drunken trees
• trees displaced from their normal vertical
alignment when permafrost melts
- Can be sign of climate warming, up to 7-8% of trees in Alaska show signs of this now.

176
Q

There are 2 possible trajectories that earth can take… Describe:

  1. The stable earth.
  2. The hothouse earth.
A
  1. Stable: High potential energy, little energy needed to move down hill.
  2. Hothouse: Low potential energy, a lot of energy needed to move out of this state (harder to reverse).
177
Q

Define (basic) this quality of snow cover that affects living organisms: Duration.

A

Onset and disappearance of snow cover

178
Q

One theory for the reason of more autochthonous lake input (“stuff” coming in from farther places) may be due to more run-off. How much is precipitation increasing per decade?

A

Precipitation is increasing by +17-19 mm per decade.

179
Q

What is a glacier?

A

Ice capable of motion - expansion and contraction over time.

180
Q

How long ago did this ice age occur?

- 3/5: Andean-Saharn

A

460-430 million years ago.

181
Q

Which tree does the brown __tree name__ longhorn beetle threaten (invasive sp)? Their larva eat tree phloem.

A

Brown spruce longhorn beetle – threatens spruce forests under temporary periods of stress

182
Q

Describe this layer of a nivean environment, and give an example of an animal that may live here.

  • Supranivean
A

at or above the snow surface

• Area where some animals travel e.g., polar bear, Canadian lynx

183
Q

Many trees depend on recurring fires to reproduce. Describe this type of post-fire regeneration and list some plants/trees that use this method:

  • Serotinous or semiserotinous cones.
A
  • Serotinous or semiserotinous cones (e.g. jack pine, black spruce)
  • Jack pine cones remain closed until the heat of a fire opens them to release the seeds inside (75% of cones)
  • Why would serotinous cones require heat of fire to open? It protects them from rain, frost, bacteria, fungus, most rodent teeth.
184
Q

What caused the formation of peatlands in Canada? What type of plants first grew?

A

Retreating glaciers left holes created by melting blocks of ice
• Water could not drain into the underlying rock strata because of permafrost
• The layer of cold water was colonized by sphagnum moss
• Over time, develops a floating mat of vegetation
• may support shrubs and small trees.

185
Q

Describe the formation of sea ice in terms of:

- What does formation begin with (___ crystals, where?)

A

Begins with frazil ice crystals (3-4mm in diameter)
• At surface – begin accumulating and bonding with
one another, chunks consolidate.

186
Q
  • Reading by Warwick *

In terms of the ecological implications of changes in the arctic cryosphere, what effect does the warming arctic cryosphere have on: Sea ice.

  • Animals (polar bears, walruses), water mixing + productivity, how it’ll change human behaviour.
A
  • Shift from birds and mammals to plankton and pelagic fish.
  • Decline in condition and reproductive success of polar bears.
  • More photosynthesis but less mixing means less nutrient distribution in deeper layers, limiting phytoplankton productivity.
  • Changes in food webs.
  • Increases in shipping, tourism, and oil/ gas/ mineral exploration that will further damage the habitats of animals.
187
Q

In terms of infrared radiation, describe good emitters/good absorbers, compared to a poor absorber..

A
  • Good emitters are also good absorbers, and good absorbers reflect little energy (ie. dark pavement).
  • Poor absorbers reflect a lot of energy (ie. snow won’t melt in sunshine).
188
Q

Impacts of disturbances are influenced by spatial factors, temporal, and magnitude.

Describe spatial factors (4).

A
  • Size
  • Shape
  • Connectivity
  • Proximity to unaffected areas
189
Q

In North America, what kind of forests would you find southwest?

A

Aspen parkland and prairie grasslands.

190
Q

Define: Acclimation.

A

Acclimation:
• Non-heritable modification of characters
• Caused by exposure to environment
• Changes readily reversible

191
Q

There are 3 zones of the boreal forest, describe:
- Forest-tundra ecotone

in terms of: where it’s found (global? Canada?)

A
  • wide across much of northern Canada, but discontinuous patches across Alaska
  • especially wide in central Siberia– Russians often treat it as a separate biome, not a transition between 2 biomes
192
Q

Increased precipitation causes increased dissolved organic carbons in lakes, = increased thermal stratification and changes in transparency. How does this influence lakes?

A
  • lead to shallower thermoclines and reductions of productive habitat in littoral zone ↳ less light can hit deeper spots.
  • Also less light which helps plant growth
193
Q

Ice melts when the temperature rises to 0°C – the melting point. Freezing is different – needs an ice catalyst (dust particle, impurity).

At what temperature does pure water solidify at?

A

-40C at standard pressure.

194
Q

Landfast first-year sea ice has a surface ___ layer and bottom ____ ice layer.

A

Landfast first-year sea ice has a surface frazil layer and bottom columnar ice layer.

195
Q

Describe Ephemeral snow in terms of:

  • General description
  • Snow depth
  • Layers of snow
A

Thin, warm, wet, lots of melt

  • Snow depth: 0-50
  • Layers of snow: 1-3
196
Q

What is happening with algae and their O2/CO2 use/release during these stages from winter to summer?

  1. Pre bloom:
  2. Increased sunlight, algal bloom:
  3. Maximum sunlight, ice melting (=melt ponds):
A
  1. Pre bloom (net heterotrophic): Using oxygen, making CO2.
  2. Increased sunlight, algal bloom: Net O2 production, using CO2.
  3. Maximum sunlight, ice melting (=melt ponds): Too much biomass, nutrients limited. Ice melt and too much light determine bloom end, the algae falls to the bottom and feeds other benthic organisms.
197
Q

Nutrients are largely a function of brine salinity in early spring. However, nutrient demand by blooming algae can rapidly surpass nutrients available in the sea ice.

What 2 things can replenish nutrients?

A

Desalination and under-ice turbulence can replenish sea ice nutrients from the water column.

198
Q

What is refugia?

A
  • Areas that provide suitable habitat for an isolated population of a once more widespread species; have not undergone ecological changes.
199
Q

Describe how lakes and rivers type can influence the amount of frozen ground present.

A
  • sources of heat in colder areas
  • Water is warmer than the surrounding air, so keeps the ground beneath warmer in the winter
  • No frozen ground beneath
200
Q

What proportion of Ontario forest disturbances are caused by:

  1. Blowdown and weather damage:
  2. Harvesting:
  3. Fire area:
A
  1. Blowdown and weather damage: 44%
  2. Harvesting: 30%
  3. Fire area: 26%

*know in general, not exact numbers

201
Q

Define: Mycorrhizae.

A

Most taiga plants have mycorrhizae – interweavings of fungal mycelium and root tissue.

Mycorrhizae are a mutualism between plants and fungi.
• the plant provides the fungus with carbohydrates
• the fungus helps the plant get nutrients from the soil, like nitrogen

202
Q

What 4 ways can endotherms lose heat (modes of heat transfer), and to what is the heat transferred to?

A
  1. Evaporation: Lose moisture.
  2. Radiation (+ or -): Heat waves.
  3. Condition (+ or -): To objects.
  4. Convection (+ or -): To air
203
Q

The functioning of Boreal lakes is tightly linked to Boreal forest through terrestrial dissolved organic carbon (DOC). What are some sources for this DOC?

How does this influence the lake?

A
  • Decomposition of leaves and other plant material
  • Autochthonous and allocthonous input of energy (sediments or other things that came from other places)
  • Affects physical characteristics of the lake
  • Slightly “stain” the water clarity
  • Attenuates light
  • Affects thermal stratification
204
Q

Define: Palsas.

Size?

Grow in, below, or above the active layer?

A

Low frost heaves that contain permanently frozen ice lenses (larger, 1-3+ m).
Palsas grow in the active layer.

205
Q

Define: Global carbon cycle (basic).

A

movement of carbon, in its many forms, between the atmosphere, oceans, biosphere, & geosphere

206
Q

What is CO2 fertilization?

A

The CO2 will fertilize the plants and increase food production (larger crops).

207
Q

What is secondary (ecological) succession?

A

Secondary succession takes place after an ecological disturbance

208
Q

The weight of ice sheets can cause the Earth’s surface to sink. How does it rebound, and what is this called?

A

Isostatic Rebound.
- When the ice sheets melt, the load on the crust and mantle is decreased. The mantle rebounds, and pushes up the earth’s crust until it’s back at equilibrium level.

209
Q

Define: Ombrotrophic.

A

Bogs are ombrotrophic (“cloud- fed”) – receive all of their water and nutrients from precipitation.

210
Q

Describe this principle force acting on sea ice:

  • Sea Surface Tilt
A

differences in surface level
• high and low regions caused by small differences in
gravity

211
Q

Arctic Ocean net primary productivity has increased by __% since 1998.

A

Arctic Ocean net primary productivity has increased by 30% since 1998.

212
Q

Conifers dominate the taiga for many reasons, describe how tolerating nutrient poor soil contributes. How do they make it harder for other plants to grow?

A
  • Live in immature soils that have little time to acquire organic material since the last glaciation (15k years ago).
    Fire disturbance every 1-2 centuries
    • Conifer needles are acidic. Acids drip from them during rain, slows decomposition, leach nutrients and minerals deeper into soil.
    • Shady year-round under closed-canopy – intercepts snow, which otherwise would form insulating cover on the ground.
    • permafrost develops more readily, further reducing the soil temperature.

Soils under conifer stands are cold, with slow decomposition

213
Q

Define: Fire regime.

A

Fire regime: pattern, frequency & intensity of wildfires that prevail in an area.

214
Q
  • Reading by Carlson + Windsor*

- What causes deglaciation?

A

Changes in earth’s orbit around the sun (increased energy) in addition to increased atmospheric carbon dioxide and other greenhouse gases.

215
Q

What kind of tree species would you find in the taiga (common names)?

A
  • Spruces dominate.
  • Pines are very plentiful too.
  • Balsam fir, poplar, birch, tamarack.
216
Q

Jack pines have serotinous cones that require fire heat to open. Why is this beneficial?

A

It protects them from rain, frost, bacteria, fungus, most rodent teeth.

217
Q

What kind of disturbances are mouse reliant on?

A

Moose are a fire-dependent species
• needs fire to create burnt-over habitat rich in deciduous
browse, which will support these giant herbivores for a decade or two.

Moose more likely in areas that had been burned or logged - They prefer disturbed areas! Mouse populations are declining in southeastern MB.

218
Q

Disturbance ecology shifts the focus from successional development of indefinite _____ communities to the dynamics of ecosystems (i.e. things keep changing).

A

Disturbance ecology shifts the focus from successional development of indefinite equilibrium communities to the dynamics of ecosystems (i.e. things keep changing).

219
Q

What is sphagnum? Is it common?

A
  • Sphagnum – responsible for most of the organic buildup in the extensive peatlands of the north.
  • Sphagnum covers more of the world’s land surface than any other single plant genus.
  • between 100 and 300 species of Sphagnum in the world (~50 sp in Canada)
220
Q

Define: Weather:

A
  • Day to day state of the earth’s surface conditions.
221
Q

What is basically occurring in an ice age?

A

A long term drop in global temperatures, causing an extension of continental ice sheets. They grow and decay regularly.

222
Q

Why might spring be a major fire season?

A
  • Trees experience water stress in spring (lose moisture due to warm temperatures, gain little from frozen soil).
  • Usually low evapotranspiration because of short growing season, cool nights, frequent cloud cover, & sun’s low angle of incidence.
  • However, the fragile water balance can be disrupted in years with thick ice on lakes and a large high-pressure system lodged overhead
    •Under drought conditions, conifers are dry; lichen cattails, grasses & sedges are flammable
    • Fire season develops and large tracts of forest burn
223
Q

Define: Sporadic permafrost.

A

mostly in peatlands & under 10-50% of land in a defined area

224
Q

In boreal forests of Canada, insect-caused timber losses (tree death) may be up to 2 times greater than the mean annual losses due to fires. What 3 parts of trees may insects target?

A
  • bark and wood-boring beetles - can recycle dying trees
  • defoliating insects (often Lepidoptera)
  • insects that attack roots and cones
225
Q

Has carbon energy pathway to benthic fauna changed over time? Explain this by answering the following question using the Atlantic walrus (a bioindicator).

These walruses are..
• Primarily benthivores
• Can consume 57kg of bivalves per day
• Central place foragers
• Haul-out in large aggregations

1) Has there been a change to phytoplankton-derived carbon to benthos?

A
  • Lipid biomarkers used to calculate proportion of sea ice algal carbon relative to phytoplankton carbon in diet.
  • 15% decrease in sea ice concentration and 50% higher contribution of carbon from phytoplankton over time.
  • This caused 13C (carbon, not celsius) depleted compared to before when it was 13C enriched due to ice algae.
  • Increased contribution of phytoplanktonic carbon to benthos has dramatically increased over 34-years (and will amplify with continued climate change).
226
Q

Do bigger or smaller animals have a higher total metabolism?

A

larger animals.

227
Q

There are 3 types of fires, surface, crown, and ground fires.

What are passive crown fires?

A

Passive crown fires: burn along ground, then spread onto each tree
• the flame length (heat transfer) is enough to ignite individual tree crowns, but the canopy fuels and/or rate of spread are insufficient to maintain the fire both on the surface and in the crowns (active crown fire).

228
Q

How does low frequency disturbance affect biodiversity?

A

• short-lived species are outcompeted by
long-lived species
• Long-lived tend to dominate the
community

229
Q

Define: Eskers.

A

Sedimentary material (gravel or sand) despoiled by streams flowing under or through glacial ice.

230
Q

Is fire disturbance important on the tundra? Are fires here common or rare, and will this change?

A

Historically, fires were rare on tundra
• projected to increase with the warming climate,
releasing more carbon, more chance of fires.

231
Q

___ caused timber loss may be 2x greater than the loss from fires.

A

insect

232
Q

The boreal forest has 3 zones based on tree cover, are they classified by specific species or something else?

A

Vegetation types are defined by community structure

• not specific species (which may be interchangeable)

233
Q

Do alpine tundras have permafrost? Do their soils drain?

A

Alpine tundra typically does not have permafrost

• soils are generally better drained than Arctic soils

234
Q

Is climate or weather more predictable? Is this changing in the arctic?

A
  • Climate is more predictable, weather is not.

- Both are becoming more unpredictable in the Arctic.

235
Q

In terms of ice ages, what are “interglacial”?

A

Pulses of warm periods.

236
Q

Thermal gradient between body core & environment can range up to 100oC. As this gradient increases, the cost of maintaining constant internal conditions ____.

A

Thermal gradient between body core & environment can range up to 100oC. As this gradient increases, the cost of maintaining constant internal conditions increases.

237
Q

Give an example of something that can be an ice nuclei.

A

Many chemicals, contaminants, and impurities will act as ice nuclei
• dust particles, bacteria

238
Q

What was the earth called during the cryogenian ice age, and what arose from this?

A
  • “Snowball earth”.

- A lot of phyla (the first ancestor) arose.

239
Q

Are forests with periodic surface fires less flammable? Why or why not?

A

Forest with periodic surface fires much less flammable than one where light fuels have accumulated for years

  • Root systems of shrubs and trees usually left alive
  • Within 3-4 years, few visible signs left of the fire
240
Q

Describe this type of lake: Deep oligotrophic.

A
3. Deep oligotrophic lakes
• Most common
• Less nutrients
• Rocky, sandy bottoms
• Less algae
• Oxygen is found at higher levels throughout water column – best water
quality
• Thermal stratification in the summer
241
Q

What are the implications of sea ice having a lammellar (spiky) bottom?

*maybe not important

A
  • Constitutionally supercooled layer, increases surface area for algal growth and traps brine throughout the ice.
242
Q

What is the issue with increased CO2 in the atmosphere?

A

CO2 increases the atmosphere’s ability to hold heat

243
Q

what 3 things must plants & animals deal with in cold boreal conditions?
*not vital

A
  • extended periods of cold • Dry and drought

* Low nutrients

244
Q

In North America, what kind of forests would you find southeast?

A

Deciduous hardwood forests.

245
Q

What TYPE of tree species would you find in the taiga (ie. conifers, deciduous, etc)?

A

Floristically poor, relatively few species but mostly conifers, some deciduous trees along water ways.

246
Q

Define: Adaption.

A

Adaptation:
• Accumulation of genetic changes
• Increases survival or reproduction in a particular environment
• Not readily reversible

247
Q

Eutrophication in lakes can deplete oxygen at the bottom of lakes and thus kill some primary producers, how does this impact predators (they also lose visual acuity in murky waters)?

A
  • Less primary consumers
248
Q

Describe how soil type can influence the amount of frozen ground present.

A

Soils freeze differently (i.e. light- colored soils freeze sooner and stay frozen longer than dark)
• light colours reflect energy, dark absorbs
• Pavement versus snow

249
Q

How do different isotopes of carbon alter the fractionation in kinetic and chemical reactions?

A
  • Diffuses at a slower speed

* Harder to break chemical bonds

250
Q

Qualities of snow cover influenced primarily by ___ and the occurrence of ___ ___ or ___-____ cycles

A

Qualities of snow cover influenced primarily by wind and the occurrence of winter thaws or freeze-thaw cycles

251
Q

Describe why this hypothesis for extinctions may not be true: Introduced species - “overkill” by human hunters.

A
  • People entered North America via the Bering land bridge.

- Problem: Very few humans to wipe out so many species.

252
Q

Defoliation can be caused by ______ which feed voraciously on new foliage & defoliate trees at high densities

A

jack pine budworm

253
Q

How does a glacier form? What is the firn limit?

A
  • Begins with snow (called firn- granular snow that’s not yet compressed) that’s modified through geological processes.
  • Under the firn limit is ice, above it is snow.
  • Glacier size depends on the balance of accumulation vs ablation.
254
Q

Acclimation vs adaption

A

Acclimation:
• Non-heritable modification of characters
• Caused by exposure to environment
• Changes readily reversible

Adaptation:
• Accumulation of genetic changes
• Increases survival or reproduction in a particular environment
• Not readily reversible

255
Q

What 2 things happen to plant tissues when water freezes?

A
  • destroys living tissue due to the expansion of water inside the cell
  • sharp ice crystals can damage and kill cells
256
Q

How are bogs “fed”? Describe this. How common are they, and what “type” are they?

A

Bogs are ombrotrophic (“cloud- fed”) – receive all of their water and nutrients from precipitation.

  • All bogs are peatlands and bogs are the most widespread peatlands in Canada (followed by fens).
257
Q

Radiant energy from the sun warms the earth’s atmosphere and surface (water and soil) and becomes heat energy. Describe this mechanism by which energy is transferred through the atmosphere:

  • Convection
A
  • Movement of a fluid (water or air).
  • Transfer of heat (current) from one place to another.
  • Radiator heats air, it becomes less dense and it rises.
258
Q

Is cryosphere (glaciers and ice) a biome?

A
  • Typical geography and biology textbooks describe tundra as the coldest biome
  • Permanent ice as biome is a novel concept
259
Q

Boreal forests recover quite fast from fires (1-2 years in some cases), and fires can cause a habitat mosaic. What does this mean?

A

• More diverse habitats can elevate species diversity in an area.

260
Q

Phytoplankton blooms happen during and after sea ice breakup (after ice algae blooms).

• How is a warming climate changing this phenology?

A
  • Sea ice is breaking up earlier, causing earlier algal blooms and larger phytoplankton booms, followed by periods with no bloom (before ice returns).
  • There is increasing water thermal stratification due to climate change too.
261
Q

Is sea ice disappearing in a linear or non linear rate? What are the consequences of this?

A
  • Ice minimum extent is decreasing at a non-linear rate
  • Ice melt contributing to freshening of surface water = greater stratification
  • more ocean is exposed every year = more light and increased atmosphere-ocean coupling.
262
Q

What do fire cycles determine? What’s the time scale for average natural fire cycles, and does it differ in some areas?

A

Fire cycle – determines many of the plant dynamics • Natural fire cycle averages 50-200 years
• Fire-prone areas (sandy ridges, rocky outcroppings) burn more frequently.

263
Q

Radiant energy from the sun warms the earth’s atmosphere and surface (water and soil) and becomes _____ _____.

A

heat energy.

264
Q

Frozen water expands __% in volume.

A

Frozen water expands 9% in volume.

265
Q

There are 3 zones of the boreal forest, describe:
- Open boreal forest (northern)

in terms of: canopy (and result of that), another name, and why it may be given this name

A
  • Trees grow far enough apart that a canopy never forms – sunlight & snow penetrate
  • Also called the lichen woodland: open woodland, with a sparse stand of conifers set in a ground layer dominated by lichens
266
Q

Many trees depend on recurring fires to reproduce. List 3 types of post fire reproduction.

A
  1. Light, wind dispersed seeds.
  2. Serotinous or semiserotinous cones.
  3. Stump sprouting.
267
Q

Do jack pines encourage or prevent fires in the boreal forest? How?

A

Jack pine: self pruning and self-shedding reduce risk of crown fire

  • Sheds most of its dead branches and needles to reduce buildup of combustible fuels in the canopy
  • Self-pruning low on the trunk removes ladder branches (which can carry a surface fire up into the crown)
268
Q

Describe this area within the boreal zone in terms of trees (basic): Tundra.

A

land without trees

269
Q

Taiga & tundra are the primary terrestrial biomes in the North
• Why not in the South?

A

There’s much more land mass in the north, the south is mostly water.

270
Q
  • If more carbon enters a pool than leaves it, that pool is considered a net ____ ____.
  • If more carbon leaves a pool than enters it, that pool is considered net ____ _____.
A
  • If more carbon enters a pool than leaves it, that pool is considered a net carbon sink.
  • If more carbon leaves a pool than enters it, that pool is considered net carbon source.
271
Q

Why are black spruce’s considered fuel ladders?

A
  • Stand structure and tree architecture of black spruce are highly conducive to fast moving crown fires
  • Thick moss-covered forest floors readily transfer flames to the lower limbs and into the canopy
272
Q

Describe this Milankovitch cycle explaining changes in insolation reaching earth:
- Eccentricity.

  • How does it vary, and how long is the cycle?
A
  • Varies from circular to elliptical.

- 100k year cycle.

273
Q

Describe the formation of sea ice in terms of:

- What temperature does it form at?

A

Forms at ~ -1.8°C

274
Q

How can we know that temperature changes have occurred in northern Manitoba?

A
  • Can identify shifts in northern boundaries of forests, due to pollen records of bogs, aspen, and spruce reflect tree line shifts over time (indicating cooling and warming periods).
275
Q

INA bacteria presence on plant surfaces causes ice nucleation at temperatures 5-8° warmer than in the absence of the INA bacteria. How have these bacteria been modified and why? What is it called?

A

Ice-minus bacteria: genetically modified mutant that doesn’t produce the ice-nucleating surface protein.
•Prevents crop damage

276
Q

How do algae (which are mostly benthic adapted, like to stick to surfaces) get into sea ice? 2 main methods.

A
  1. Scavenging: Picked up by fazil crystals (which form at depths), then get stuck to the ice once it forms.
  2. Sieving: Turbulance or brine drainage causes the sticky algal cells (due to exopolymeric substances/EPS).
277
Q

What is primary (ecological) succession?

A

Primary succession

takes place on land where no organic soil exists – pioneer plants

278
Q

What’s the difference between a poikilotherm and ectotherm?

A

Poikilotherms: animals that have a variable body temperature over time.
e.g., moths and butterflies

Ectotherms: generally do not produce sufficient heat to maintain a constant body temperature. Rely primarily on external sources of heat.
– However, may have constant temperature depending on behavior and environment.

279
Q

Rising global temperatures may increase the number of lightning strikes over the rest of the 21st century

• __% increase for each degree Celsius

A

Rising global temperatures may increase the number of lightning strikes over the rest of the 21st century

• 12% increase for each degree Celsius

280
Q

What does “stability” mean in the context of earth?

A

More likely to stay in the current state.

281
Q

Many trees depend on recurring fires to reproduce. Describe this type of post-fire regeneration and list some plants/trees that use this method:

  • Light, wind dispersed seeds.
A
  • Light, wind-dispersed seeds (e.g. birches, poplars)

- Aspen catkins produce small fruit that split to release lots of tiny, cottony seeds that are dispersed by the wind

282
Q

Arctic temperatures are changing __ -__ times faster than in other areas.

A

3-4x

283
Q

How many terrestrial mammal species went extinct in the late Pleistocene?

A
  • 35-40 large bodied mammals (mammoths, giant ground sloths, sober tooth cats, giant bisons).
284
Q

Which area of the boreal forest has the most fires: east, middle/central, or west?

A

Mostly in the middle, and they go quite far up north.

285
Q

T or F: Old arctic ice is disappearing.

A

True - Ice is becoming younger and thinner.

286
Q

List the 2 sources of changes in thermal energy that can cause ice ages.

A
  1. Earth centre (inner core).

2. Insolation (from the sun).

287
Q

Vegetation patterns exist in the tundra. Describe: Fell fields

A

Fell fields – stony tundra area where vegetation is thin and discontinuous – supplants heath tundra and tussock tundra in drier areas

288
Q

How often do ice ages happen?

A

They are cyclical, 44k - 110k years.

289
Q

does brain size correlate with trophic position?

A

yes

290
Q

In North America, what kind of forests would you find west?

A
  • Montane and coniferous, and moist coniferous closer to the pacific coast.
291
Q

Rising global temperatures may increase the number of lightning strikes over the rest of the 21st century - if there’s a 4C projected increase in temperature between 2014 and 2100, how many more lighting strikes will there be (%)?

A

48% more lightening strikes.

292
Q

What is the 1st and 2nd largest component of the climate system?

A

First is oceans, the second is cryosphere.

293
Q

Describe this type of lake: Eutrophic.

A

1) Eutrophic
• Shallow and have murky water and mucky, soft bottoms.
• Nutrients carried into lake from landscape
• High levels of biological productivity
• High biomass of algae, fish and other aquatic organisms
• Fine balance – too much algal blooms and eutrophication (anoxic zones)

294
Q

Are there exceptions to Bergmann’s rule? Why?

A
  • Yes, animals tend to be smaller at higher latitudes.

- Likely attributed to primary productivity, if you’re bigger, you need more energy.

295
Q

Define: Kame.

A

A hill or hummock composed of stratified sand and gravel laid down by glacial meltwater.
- Accumulates in a depression on a retreating glacier, and is then deposited on the land surface with further melting.

296
Q

Describe how snow can influence the amount of frozen ground present.

A

A thick layer of snow acts like a thermal blanket – traps heat

297
Q

What 3 potentially problematic things can be released with thawing permafrost?

A

1/2. Releases CO2 and methane (microbial decomposition; speeds up climate warming).
3. Mercury (15 million gallons, 10x more than the amount of Hg emissions over the last 30 years).

298
Q

Conifers dominate the taiga for many reasons, describe how their low surface area contributes contributes.

A
  • Needleleaf - narrowness reduces surface area through which water may be lost (transpired).
  • Needles also have thick waxy coatings - a waterproof cuticle
  • Stomata are sunken and protected from drying winds.
299
Q

What are isotopes of carbon? List the 3 main ones, and what makes them an isotope. Which ones are more common?

A
  • 14C (radioactive), 13C and 12C (stable isotopes),
  • Forms of the same element that differ in the amount of neutrons in the nucleus.
  • Lighter ones are most common.
300
Q

List 4 qualities of snow cover that affect living organisms.

A
  1. Duration.
  2. Thickness.
  3. Hardness.
  4. Density.
301
Q

What 2 things cause seasons?

A
  1. The tilt of earth on the N/S rotation axis.

2. It’s orbit around the sun.

302
Q

Why is sphagnum able to dominate?

A

Dominance of Sphagnum due to ability to absorb and retain water.
• Absorbs 8 times its weight in water due to leaf anatomy
• Closely packed sphagnum creates wick-like columns of moss, which lift water to the top of a sphagnum hummock (capillary action).

303
Q

Lake trouts prefer temperatures of <13C and high dissolved oxygen concentrations, meaning they have a much smaller possible habitat in summer. Have the thermal and dissolved oxygen barrier changed over time? I.e. have summer gotten longer and hotter?

A

Evidence of earlier ice melting, longer springs, and the cooling of lakes to <15°C occurring later in the fall… suggests that the summer period has shifted later into the year.

Due to the later onset of fall, ice-on became delayed, and paired with earlier ice-offs, the winter ice-covered period became shorter over time.

304
Q

Conifers dominate the taiga for many reasons, describe how their conical shape and dark colour contributes.

A
  • Conical shape – promotes shedding of snow & prevents loss of branches.
  • Dark color – helps the foliage absorb maximum heat from the sun and begin photosynthesis as early as possible
  • start of the growing season
305
Q

Describe how (domed) bogs are established, including the role of sphagnum (6 major steps).

A

Sphagnum can establish whenever precipitation > evaporation
• Water fills depressions
1. Spreads mostly by spores (it’s a moss!)
2. Over many years grow into a large mass of floating moss
• “floating blob” stage
3. Starts diverting water – blocks flow into and out of the site
• Cuts off nutrients & minerals necessary for competitors
• Accumulation seals off the mineral soil
4. Begins to produce acid, killing off plants that can’t tolerate it
5. High acid and low oxygen kills many decomposers – dead plant material starts to accumulate
6. Mounds of organic material begin to collect. Peat accumulates, bog takes on a raised (domed) profile

306
Q

What are some ways animals rely on snow and sea ice?

A

Protection from predation Thermoregulation

Nursing, birthing and resting

307
Q

How does arctic cod avoid freezing?

A

Arctic cod, use antifreeze glycoproteins to lower the freezing point of their tissues, blood and digestive fluids

308
Q

After sphagnum establishment – some species perform counter tactics. List 3 species that can do this, and how.

A
  • Leatherleaf, Labrador tea, bog rosemary.

Leathery, sturdy, evergreen leaves remain on plant for several years
•less energy & nutrients needed to start growth in spring
•many fine, hairlike structures on undersurface of leaves → to soak up more water
•thick, waxy coat on upper surfaces – reduces water loss

309
Q

How is climate change influencing benthic organisms (especially macroalgae, is there more or less of it now)?

A

There’s much more macroalgae cover.

310
Q

Define: Insolation.

A

Total amount of solar radiation energy received on a given surface area during a given time.

311
Q

Describe the Wisconsinan glaciation (most recent in North America) in terms of:

  • How long ago it was.
  • How much of land was covered in ice.
A
  • 85,000 - 11,000 years ago.

- Covered 80% of North America.

312
Q

Describe the formation of sea ice in terms of:

- What does it become in calm waters vs windy waters?

A
  • In calm waters – grease ice

* Windy waters - pancake ice

313
Q
  • Reading by Dredge*

- How long ago did the last ice sheet disappear from Churchill, and what was the name of this ice sheet?

A

The Laurentide Ice Sheet disappeared from Churchill 8000 years ago.

314
Q

Vegetation patterns exist in the tundra. Describe: Wet tundra

A

Wet tundra: standing water remains over most of the summer • more widespread and less uniform than tussock tundra
• develops in flat, marshy areas, usually with Cottongrass
plus true sedges

315
Q

Many trees depend on recurring fires to reproduce. Describe this type of post-fire regeneration and list some plants/trees that use this method:

  • Stump sprouting.
A
  • Stump sprouting (aspen, paper birch).

- Birch & aspen regenerate vegetatively by shoots and suckers arising along the long lateral roots.

316
Q

Does tree line advancement northward depend on temperature?

A

“Water availability rather than temperature appears to be an increasingly limiting factor for the northward progression of the treeline”
Advancement will be patchy where there’s enough
moisture

317
Q

How can wind cause forest disturbances?

How many hectares can be destroyed by this?

A

Severe weather events can cause small or large areas of forest to blowdown
• can result in some or all trees within an area to break off (windsnap) or uproot (windthrow)

Blowdown events usually range from 5 to 50 hectares in Ontario
• up to 500,000 hectares

318
Q

Snow depth/ melt pond coverage largely control light availability in sea ice.

How does albedo differ between snow and ice?

A

Albedo
̈ Snow – 0.8
̈ Ice – 0.4

319
Q

What is the result of glacier abrasion, and how does it occur?

A

Smoothing of rocks as a result of ice flow.

320
Q

Define: Peat.

Where does peat form?

A

Peat: an accumulation of partially decayed vegetation • forms in wetlands or peatlands, variously called bogs, moors, muskegs, pocosins, mires, and peat swamp forests.

321
Q

What is a cryosphere?

A

Places were water is in solid form (not solely polar):

  • Frozen ground
  • Glaciers
  • Icebergs
  • Snow cover
  • Sea ice
  • Found every continent (even Africa, snow topped mountains)
322
Q

Pure water solidifies at -40C. What does water require to freeze at lower temperatures, and at what temp can it freeze at with these things?

A

Above -40°C, pure water needs a catalyst to transition to ice.
• Ice crystals can act as catalysts.
• the freezing point of water under equilibrium conditions
(when an ice crystal is present) is 0°C.

323
Q

What are thermokarst lakes, and how does permafrost make them form?

A

• Irregular surfaces of marshy hollows and small hummocks • Caused when permafrost thaws and soil collapses

324
Q
  • Reading by Warwick *

In terms of the ecological implications of changes in the arctic cryosphere, what effect does the warming arctic cryosphere have on: Snow.

  • Animals, microbial life, plants.
A
  • Increasing winter precipitation in the arctic.
  • Decreased snow cover duration (due to warmer air temperatures).
  • This together is causing winter rains and freezings that cover the surface of snow in ice, impeding animal foraging.
  • Organisms in snow (bacteria, fungi, snow algae) provide organic carbon and nutrients to the soil under snow during melts, and warmer temperatures can cause southern plant life to invade and outcompete the lichens and bryophytes present.
  • Without an insulating layer of snow, buds and shoots can be damaged and killed, reducing primary production.
  • Changes in growing season change the food resources available to arctic herbivores.
325
Q

Why are peatlands important? What is then a problem if peat is degrading due to climate change?

A

Peat is important because it stores organic carbon that otherwise could be released into the atmosphere as carbon dioxide or methane (carbon sink).

If peat is disappearing, there will be more CO2 and methane in the environment.

326
Q

Reading by Pruitt

What is pukak (in terms of snow), how is it made, and why is it important?

A
  • Metamorphosed layer of large snow crystals, extending 10cm or more.
  • In the absence of wind, snow crystals settle undisturbed on other snow crystals, and are metamorphosed by heat and moisture rising from the earth below.
  • Important for subnivean mammals.
327
Q

Describe how wind direction can influence the amount of frozen ground present.

A
  • If a slope faces into the wind, the ground will lose more heat through convection
  • Wind blows snow, making the ground even colder
  • Wind-blown slopes are more probable to have frozen ground.
328
Q

What is the low arctic in terms of:

  • General area (relative to tundra)
  • Vegetation, ground, amount of snow
  • Winter thaw or no?
A
  • southern part of tundra
  • complete vegetation cover
  • patches of shrubs
  • lots of snow
  • Can have winter thaw
329
Q

Describe this type of lake: Mesotrophic.

A

2) mesotrophic
• a medium amount of nutrients
• usually clear water with submerged aquatic plants
• These lakes stratify: colder bottom, warmer surface

330
Q

What are 4 big consequences of water eutrophication?

A
1. Reduce water clarity
• Affect acuity of visual predators
2. Harms water quality
• Large algal blooms
• Cyanobacteria – toxic
3. Raise water pH
• Blind chemosensory predators
4. Hypoxic and Anoxic zones at bottom • Depleted or removed O2 at bottom
due to microbial decomposition
331
Q

Describe why this hypothesis for extinctions may not be true: Glaciation-induced climate change.

A
  • Problem: Extinction were very abrupt (for geological time).
  • Only seemed to affect large mammals or birds.
332
Q

Define: Continuous permafrost.

How thick is it?

A

Continuous→ occurs under all the land surface in a defined area, except beneath large water bodies
• Often >100m thick

333
Q

T or F: It’s warmer closer to ground surface than at snow surface

A

True

334
Q

in terms of this principle force acting on sea ice, the speed is predictable - sea ice that drifts freely moves at __% of the wind speed

A

2%

335
Q
Canada is home to:
• \_\_ of World’s lakes that are > 1km^2
• \_\_ of 50 longest rivers across World 
• _00,000 km2 of surface water
• Overall ~ \_\_% of World’s wetlands
A

Home to:
• Half of World’s lakes that are > 1km2
• 5 of 50 longest rivers across World • 800,000 km2 of surface water
• Overall ~ 25% of World’s wetlands

336
Q

Fire size is a function of ____ and _____ (2 things that contribute to the size).

A

Fire size is a function of fire weather severity (e.g. dryness) and the connectivity of fuels (vegetation) on the landscape.

337
Q

Generally, Ectotherm = Poikilotherm and Endotherm = Homeotherm. But what about these 2 examples?

  • Several deep sea fish
  • Hibernating Mammals
A
  • Several deep sea fish (homeothermic ectotherms)

* Hibernating Mammals (poikilothermic endotherms)

338
Q

Lake trouts prefer temperatures of <13C and high dissolved oxygen concentrations. Why might the summer season be problematic?

A

Temperatures close to the surface are hotter, and there’s less oxygen at the bottom of the lake, meaning that it’s preferred habitat is much smaller in the summer.

339
Q

Fire is the primary agent of change in the boreal forest. Describe 2 ways it can cause change.

A
  1. releases nutrients
    • fires reduce plant litter (leaves, logs & conifer needles) on the forest floor and releases nutrients from these materials
  2. allows more sunlight
    • Fires also open the canopy to sunlight which stimulates regeneration from seeds and roots (growth and reproduction).
340
Q

If peatlands are rich in carbon, what would happen in high temperatures and fires?

Is a feedback loop established?

A
  • Higher temperatures could increase the rate of decomposition, releasing carbon.
  • Increased fire frequency could release more carbon trapped in peat.

Positive feedback between increasing temperatures and CO2 emissions
• Exacerbates the problem

341
Q

There are 3 zones of the boreal forest, describe:
- Closed boreal forest (southern)

in terms of: density, canopy, soil, nutrients, permafrost.

A
  • Dense forest, trees form a closed canopy

* Cold soils, low nutrients, permafrost

342
Q

How does high frequency disturbance affect biodiversity?

A

• no time for certain species to mature
• community is dominated by species with a
rapid growth rate, a short life span, and a strong ability to colonize disturbed areas (annuals)

343
Q

Define: Depth hoar.

A
  • Depth hoar→ coarsest grains of snow usually found near the soil surface (sugar snow)
  • Grains are larger and poorly bonded to one another
  • Forms subnivean zone → Small animals can use this as shelter in winter
  • Destabilizes snow→ avalanches
344
Q

What are the relative % surface albedos of:

  • Fresh snow or ice
  • Soil
  • Water
A
  • Fresh snow or ice: 60-90%
  • Soil: 5-30%
  • Water: 5-10%
345
Q

Reading by Pruitt

What is api snow?

A

The snow on the ground (in the taiga).

346
Q

Vegetation patterns exist in the tundra. Describe: Polar desert

A

Polar desert – barren areas of bare rock, shattered bedrock, and sterile gravel.
• An extreme form of fell field in the high Arctic
• Lichens important – key for large herbivores

347
Q

What is the active layer in permafrost?

A

Above permafrost; seasonal thawing and freezing.

348
Q

How did moving ice change geology of the Canadian Shield?

A
  • Glacier abrasion (due to ice flow) can smooth rocks.
  • Plucking (glacier picks up rocks and sediment and deposits it elsewhere).
  • Caused deposition of rich silt.
  • Some areas that previously had remnants of earth’s original crust were overlaid by sedimentary rock (limestone).
349
Q

What are some natural causes of disturbances? There are 6, list 4.

A
Natural:
• Floods
• Glaciers
• Fires
• Volcanoes
• Frost heave
• Ocean currents
350
Q

There are 3 types of fires, surface, crown, and ground fires.

Describe surface fires in terms of what conditions cause it to occur, and what are the consequences? Are there benefits?

A
  • Only occurs if moisture is moderate before the fire, and wind conditions are moderate during the fire
  • May kill thin-barked trees if it gets hot enough, but doesn’t destroy mature forest
  • Thins out forest – kills some saplings, shrubs, herbs, & weaker trees
  • reduces combustible fuels, decreasing chance of future fires
351
Q

Benthic-pelagic coupling is crucial for energy cycling.

Ice algal primary production contributes __ - __% depending on area.

A

Ice algal primary production contributes 3 - 60% depending on area.

352
Q

List the 2 hypothesis for extinction in the Pleistocene (it was likely a combination of both causing it).

A
  1. Glaciation-induced climate change.

2. Introduced species - “overkill” by human hunters.

353
Q

Describe this quality of snow cover that affects living organisms: Thickness.

How does it vary?

A

How deep the snow is – critical habitat
for small mammals
• Varies by latitude, regional climate, moisture, elevation

354
Q
  • Reading by Carlson + Windsor*

- How does glacial melting influence sea level?

A

Ice melting can cause rapid oscillations (both increases and decreases) in sea level.

355
Q

Describe this Milankovitch cycle explaining changes in insolation reaching earth:
- Precession.

  • How does it vary, and how long is the cycle?
A
  • Earth wobbles as it rotates, causes different North Star.

- 26k period.

356
Q

How does a low-severity fire encourage or prevent further fires? How does it influence other tree/ plant life?

A
  • May kill most of the vegetation but removed little of the standing biomass…
  • The high fuel load now increases the risk of another more severe fire.
  • Thick (> 1 meter) mats of sphagnum moss can protect the soil from high severity burns
  • The resulting postfire forest floor mosaic limits the recolonization of black spruce seedlings to areas of highest severity and shallowest remaining organic layer
357
Q

What are some human causes of disturbances? There are 6, list 4.

A
Human-caused:
• Mining
• Agriculture
• Logging
• Urbanization
• Road construction 
• Ecotourism
358
Q

How can boreal forest “seed banks” promote growth?

A

Some understory plants survive as seeds buried in the soil for decades only germinate when canopy & leaf litter removed by fire.

359
Q

Once polar darkness sets in, autotrophic community members must employ survival tactics. How do sea ice diatoms survive?

A

Sea ice diatoms likely use a combination of facultative heterotrophy, a reduced metabolic state, and utilization of intracellular energy stores as well as previously released exopolymeric substances.

360
Q

What is the length of fire cycles controlled by? In boreal forests, are older trees more or less flammable?

A
  • Length of the cycle controlled by moisture balance
  • Precipitation vs. evapotranspiration
  • Most boreal forests equally flammable regardless of age (after first decade)
361
Q

How long ago did this ice age occur?

- 5/5: Quatenary

A

2.6 million years ago to present.

362
Q

What are hyaline cells in sphagnum? What does this cause (that’s potentially harmful)?

A

Hyaline cells – water-storage facilities
Also responsible for acidification – inner surfaces are lined with positively charged hydrogen ions
• released whenever another positive ion bonds in its place (creates acid)
- This ion exchange removes nutrients, acidifies the site, and inhibits decomposition.
- The pH of the water of Sphagnum bog may range from 3.7 to 4.9 (Parihar 1965)

363
Q

How big of an area do black and white spruce span in North America, and what layer (+depth) can they grow in? Do they dominate? Do they grow in wet or dry areas (drainage)?

A

Black spruce and white spruce are transcontinental – Very shallow root systems, can grow in an active layer only 25 cm thick. Dominate northernmost part, south of treeline.

Black spruce grow in poorly drained low lying areas, while the white spruce is in well-drained, dry uplands.

364
Q

Do bogs or fens have more plant species? Which one is more common?

A

The total number of plant species is greater in fens

than in bogs – bogs are more widespread

365
Q

What does the emerald ash borer do to trees? Can it be eradicated? How many trees could it kill in the next 10 years?

A

Beetle that girdles tree – removes bark
- Nothing can be done once infested
- First found in Windsor, ON in 2002
- Winnipeg – fall 2017
Could kill up to 350,000 ash trees in next 10 years
-Injected with insecticide to slow infestation

366
Q

Vegetation patterns exist in the tundra. Describe: Polar steppe

A

Polar steppe – similar to fell field, but on dry fine-grained soil, silt or sand (not gravel, rock or rubble). Vegetation more complete; most plants herbaceous
• Dry, turfy appearance.

367
Q

How can movements from refugia be mapped?

A

Pollen records - pollen grains washed or blown into lakes acclimate in sediments and provide a record of past vegetation (reflecting climate conditions favourable for that vegetation).

368
Q

To stay alive in cold, life has a few options: Stay above 0C, or drop below 0C.

In terms of staying above 0C, describe migration.

A

Go to warmth and food
• Large mammals
• Birds
• Monarch Butterflies

369
Q

Describe this principle force acting on sea ice:

  • Internal Ice Stress
A

measure of the compactness, or strength, of the ice – most variable
• resistance to the motion caused by the wind force
• minimal when the sea ice pack is loosely compacted and can flow freely (e.g., in summer), but can be high when the ice is compact and cannot flow
• plays an important role in the deformation of the ice and →
in the formation of some features, e.g., ridges and leads.
• strength of ice depends primarily on its thickness. Thin ice breaks apart easily under compression

370
Q

Phytoplankton blooms happen during and after sea ice breakup (after ice algae blooms).

How do the phytoplankton blooms influence other life?

• How is a warming climate changing this phenology?

A
  • That large amount of energy drives productivity at surface

* Predators and in turn, their prey follow

371
Q

Are equilibriums ever reached indefinitely?

A

There are usually ups and downs constantly (if you graphed one parameter, it would look like waves).

372
Q

Lake Winnipeg is an example of a eutrophic lake. What nutrient causes algal blooms and where does it come from?

A
  • Algal blooms in Lake Winnipeg are associated with both point source and non-point source phosphorus loading (non point source = from a distance away)
  • Large increases in blooms since 1990 are related to land-use and higher than normal precipitation associated with climate change (washes it into lake)
373
Q

How does the slope of RMR vary for tropical vs arctic animals?

A

Slopes are steeper for tropical – use a lot more energy at lower temps than Arctic mammals

374
Q

What part of trees are good for fire resistance? List 3 species with good resistance.

A
  • Tree bark is protective.

- Paper birch, black spruce, jack pine.

375
Q

In terms of boreal forest fires; Why no change in mean fire size over time?

A

recently burned areas are less likely to burn again for a long time because of the decrease in fuel load

376
Q

As the earth orbits, it stays tilted in the same direction. When it’s tilted toward the sun, it’s ____. When it’s tilted away from the sun, it’s ____.

A

summer, winter.

377
Q

Vegetation patterns exist in the tundra. Describe: Shrub tundra

A

Shrub tundra - Low Arctic, vegetated with willow, dwarf birch, and heath

378
Q

What’s the rule of 5 in terms of bulk salinity and brine volume?

A

At approximately -5 °C, bulk salinity of 5, brine volume is 5% = liquid permeable

  • the bottom of sea ice (above 5% brine) can flow and exchange water and other stuff. Below 5%, sea ice can’t move.
  • Without the movement, there’s limited access to important nutrients for primary producers.
379
Q

Constructive and destructive metamorphism occur simultaneously… which of these would occur in this scenario?

New snow – _____ tends to be more common.
Old snow – _____ tends to be more common.

A

New snow – destructive tends to be more common.

Old snow – constructive tends to be more common.

380
Q

Variety of adaptation mechanisms exist in the sea ice atmosphere. Exopolymeric Substances (EPS) produced by sea ice diatoms and bacteria are one, describe how they fit these 3 mechanisms:

  1. Extracellular
  2. Internal and Intracellular
  3. Behavioural
A
  1. Alter physical environment (can drill holes, make their own environment; EPS-clogging of brine channels can reduce brine drainage, increasing local brine salinities – increases habitat space via depression of freezing point)
  2. Support adhesion and
    locomotion
  3. Buffer and protect against extreme physicochemical conditions (defines against bacteriophages)
381
Q

Albedo is much higher in sea ice due to the numerous impurities found in sea ice (_____ growth) compared to lake ice (____ growth).

A

Albedo is much higher in sea ice due to the numerous impurities found in sea ice (lamellar growth) compared to lake ice (planar growth).

382
Q

Describe Taiga snow in terms of:

  • General description
  • Snow depth
  • Layers of snow
A

Thin-moderate, in trees, deep depth hoar

  • Snow depth: 30-120 cm
  • Layers of snow: >15
383
Q

What is a disturbance?

A

Relatively discrete event in time and space that alters the structure of populations, communities, and ecosystems and/or changes resources, substrate availability, or the physical environment.

384
Q

Ice melts when the temperature rises to __°C – the melting point.

A

Ice melts when the temperature rises to 0°C – the melting point.

385
Q

What does it mean that Canada is the “World’s Waterkeeper”?

A

Canada houses more surface water than any other country.

386
Q

Define: Growing season.

A

period in the year during which temperatures are high enough for photosynthesis

387
Q

What is benthic-pelagic coupling? Why is it important?

A

exchange of energy, mass, or nutrients between benthic and pelagic habitats – crucial for energy cycling

388
Q

How long ago did this ice age occur?

- 4/5: Karoo

A

360-260 million years ago.

389
Q
  • Reading by Warwick *

In terms of the ecological implications of changes in the arctic cryosphere, what effect does the warming arctic cryosphere have on: Glaciers and ice shelves.

  • Benthic organisms, new/ invasive /extinctions of species, human impacts (fisheries), terrestrial environment (temperature effects).
A
  • Glaciers and ice shelves provide a habitat for microbial communities and water supplies for streams, rivers, wetlands, and the coastal ocean.
  • Increased melting may stimulate benthic diversity, and open up new terrestrial habitats for microbes, and plants.
  • Once much of the water input is gone, the cold water species will likely die off, and warm-water taxa will invade.
  • Potential effects on currents, nutrient supply, marine productivity, fisheries, loss of habitats for marine birds and mammals, changes in coastal environments.
  • Can cause faster warming of terrestrial environments.
390
Q

There are 3 types of fires, surface, crown, and ground fires.

Describe crown fires in terms of what burns, and how they form.

A

Crown fires – surface fire builds up enough heat to ignite the canopy
• common in boreal forests, rare in deciduous forests
• depends on tree canopy architecture & understory vegetation

391
Q

Related to sea surface tilt, what’s a Geoid?

A

• Geoid – undulating surface results from the uneven distribution of mass within and on the surface of Earth

392
Q

Describe this area within the boreal zone in terms of trees (basic): Taiga.

A

boreal or northern coniferous forest

393
Q

Describe this layer of a nivean environment, and give an example of an animal that may live here.

  • Subnivean
A

below the snow
• On land, created in area of depth hoar or pukak
• On ocean, area where ice and snow meet
• Denning area for ringed seals

394
Q

Define: Moraine.

A

Accumulation of glacial debris at the edge of a glacier.

395
Q

Radiant energy from the sun warms the earth’s atmosphere and surface (water and soil) and becomes heat energy. List the 3 mechanisms by which energy is transferred through the atmosphere.

A
  1. Conduction.
  2. Convection.
  3. Radiation.
396
Q

Conifers dominate the taiga for many reasons, describe how their short growing season contributes.

A

• Short growing season, but even shorter for deciduous species
• Conifers photosynthesize whenever
weather is warm enough.
• Not as efficient at photosynthesis as broadleaf plants, but don’t have to be – don’t grow new leaves
• Accumulator species – hoard minerals

397
Q
  • Reading by Warwick *

In terms of the ecological implications of changes in the arctic cryosphere, what effect does the warming arctic cryosphere have on: Water.

  • Plants, lakes/water bodies, animals, fish.
A
  • Water availability is what limited northward progression of the treelike, not temperature. The tree lines will colonize where is there is enough moisture (non-uniformly), and can shift water balance, causing lakes and wetlands to dry up.
  • Snow banks are an important source of water for wetlands, and decreasing snow duration can ruin this.
  • The degradation of permafrost soils can result in the collapse, drainage, and complete disappearance of water bodies.
  • Warmer temperatures will cause rain instead of snow, which favours growth of broadleaf over conifer species, which decrease carbon storage, increase soil respiration, and decrease the timber potential of the forest.
  • Sudden rain can result in the rapid loss of protective snow, making vegetation vulnerable to freezing, and can prevent herbivores from accessing the (frozen) vegetation.
  • Changes in climate can change connectivity in bodies of water, which is problematic for migratory fish.
398
Q

Describe how peat can influence the amount of frozen ground present.

A
  • found in marshy areas, partially decomposed dead plants - ground under peat is usually colder
  • Freezes in winter so allows heat to leave the ground
  • Dries in summer, insulates ground below
  • Land rises up because of peat plateaus (flat-topped) and palsas
399
Q

In terms of ice ages, what are “glacial”?

A

Pulses of cold periods.

400
Q

Frozen water expands 9% in volume, which injures

cells in 2 ways:

A

1) by rupturing the cell membranes

2) by affecting the permeability of those membranes

401
Q

What is the subarctic in terms of:

  • How a climatologist would define the area (temp wise)
  • How a biologist would define the area (vegetation and permafrost or no frost)
A
  • Tundra-Taiga zone
  • Climatologist: mean temp. >10C for < 4 mo.
  • Biologist: both forest and permafrost (continuous & discontinuous)
402
Q

What happens when permafrost melts in….?

Tundra –

Taiga –

A

Tundra – thermokarst lakes (soil collapses when permafrost or ice wedge thaws, creating a lake/marshy hollow)

Taiga – drunken trees (trees displaced from their normal vertical alignment when permafrost melts)

403
Q

T/F - are drunken trees a sign of global warming?

A

T

404
Q

What proportion of trees in Alaska are showing signs of global warming, presenting as drunken trees?

3-4%
5-6%
7-8%
9-10%
11-12%
A

7-8%

405
Q

Why can’t trees grow in the far north?

A

The lack of summer warmth – not cold winters.
• Photochemical reactions (photosynthesis) can
only happen if plant tissues are warm enough
• Results in low growth
Leaves and roots are a necessity; wood is a luxury

406
Q

___ are the most widespread peatlands in Canada, followed by ___

A

Bogs are the most widespread peatlands in Canada, followed by fens

407
Q

The total number of plant species is greater in ___

than in ___, ___ are more widespread

A

The total number of plant species is greater in ____

than in ___, ___ are more widespread

408
Q

____ – responsible for most of the organic buildup in the extensive peatlands of the north.

A

Sphagnum – responsible for most of the organic buildup in the extensive peatlands of the north.

409
Q

Most arctic sea ice is now

A

1

410
Q

How do cold temperatures affect animals? As in, what 3 main challenges do they face?

A
  1. Food scarcity – plants die off or buried and endotherms need more food to burn for heat production.
  2. Metabolism disrupted
  3. Freezing. Ice crystals cause damage. Cells dehydrate.