Lecture 5: Visual cortex Flashcards
Where is the primary visual cortex located?
at the back of the brain in the occipital lobe
Anatomy of V1:
Magnocellular =
Layer 4Ca
Anatomy of V1
Parvocellular=
Layer 4Cb
Anatomy of V1
Koniocellular=
layer 2 and 3
Hemisphere projections
**
where do cells in layers 2, 3 and 4b project?
extra striate cortex
where do cells in layer 5 project?
midbrain structures such as superior colliculus and pons
where do cells in layer 6 project?
LGN.
- projections are highly selective
- different cells in different parts of the layers of 6 projecting to different layers in the LGN
Anatomy of V1: Ocular dominance columns
a slab of cortical tissue running perpendicular to the cortical surface, in which all binocular cells share the same degree of ocular dominance
- separation of the image from each eye in LNG maintained in V1
- layers specific to either left or right eye
Orientation columns
a slab of cortical tissue running perpendicular to the cortical surface, in which orientation selective cells share the same preferred orientation
- respond to different stimuli in the world of varying angles
- orientation column specific to different orientations
Blobs
- what layers are they in
- what are they sensitive to?
- orientation?
- how were they discovered?
- 2,3 and V1
- brightness and colour
- monocular: are not orientation specific
- using cytochrome oxidase staining
Blobs: circularly symmetrical receptive fields
Stimulation with a particular colour in the centre elicits a response, whereas stimulation of the immediate surround elicits the reverse response
V1 receptive fields
- several LGN cells combine to input on a single V1 cell
- elongated receptive field and are more selective for what stimuli activates or excites them
3 main types of orientation selective cells:
– Simple cells
– Complex cells
– Hypercomplex cells
Receptive fields in LGN
- what are they comprised of?
- how do they respond to bar of light in any orientation?
- ganglion cells and are circular
- light in centre calls to fire, light in the surround causes it to silence
Complex cell
a cell in visual cortex with a relatively high receptive field that does not contain identifiable excitatory or inhibitory zones, but is orientation specific
-responds to a bar of a certain orientation anywhere within its receptive field
-created by grouping simple cells of the same orientation
Simple cells
an orientation selective cell in visual cortex with an elongated retinal receptive field containing excitatory and inhibitory zones
Complex cells
Motion detection
- some cells in V1 respond to stimuli moving in certain direction also
- approx 25-35% of V1 cells are strongly directionally selective
Hypercomplex cells
-generated through summating a number of complex cells (v.large receptive field)
- activity dependent on length as well as orientation
- if bar extends outside of receptive field, stops firing
Topography in V1
-cells distributed across the visual cortex so that their receptive field locations form an ordered topographic map of the visual field
Cortical magnification factor
=central areas in the visual field (the fovea) have disproportionately large area of the cortex devoted to it