Lecture 5 Flashcards
Leaves
Photosynthetic organs with a manufacturing cost
Leaf shape
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A) Hight surface area for CO2 absorption (H2O loss) and light exposure, high SA:V
Leaf Shape
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B) lower SA:V , less photosynthesis but better H2O conservation
Leaf Shape
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C) Lowest SA:V, little SA for gas exchange best H2O conservation
Leaf Shapes
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D) High SA, no self shading, must be long to achieve volume = a
Leaf Layer
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-Chloroplasts are mainly located in mesophyll cells
Layer from top to bottom
- Cuticle
- Upper epidermis
- Pallsiade Mesophyll Cell
- Bundle Sheath Cell
- Xylem
- Phloem
- Lower Epidermis
- Spungy mesophyll Cells
- Guard cell
- Stoma
- Cuticle
Leaf Fibers
- made from xylem and phloem cells bundled in leaves
- sisal and henequen come from leaves of Agave spp. *Agave sisalana *has sharp spoines that may also be used a needles, “needle and thread” plant
- ropes made from sisal in Africa - can make heavy duty rugs/mats
- has thorns on the side that would be used as a needle
Surface area to volume relationship of different leaf shapes
A) High SA:V = High surface area for CO2 absorption (H2O loss) and light exposure (example: standard broad leaf)
B) Lower SA:V = less photosynthesis but better H2O conservation
C) Lowest SA:V = little SA for gas exchange, best H2O conservation
D) High SA = no self-shading, must be long to achieve volume = a (Example: needle plant)
Chloroplast structure
Chloroplasts are mainly located in mesophyll cells
Mesophyte
- Syringa vulgaris*
- plants that live in medium conditions
Example: lilacs
Xerophyte
- Nerium oleander* (Oleander)
- plants that live in extremely dry conditions
- plants of dry areas
- multiple layers of endodermis with thick cuticle - prevents loss of water molecules
- stomatal crypts
- trichomes within crypts retain moisture near stomata
Stomatal crypts
- where the stomata is sunken into the epidermis like a little cup
- can have trichomes - prevents water loss, creates drags
- reduce moisture loss
- found mostly in Xerophytes
- example: Oleander
Trichomes
DEFINNEEEEE
Hydrophyte
- Nypmphaea* (Water lily)
- stomata on upper surface, or lacking
- lots of air space to conduct O2to submerged portions of plant
- less xylem tissue
- air spaces help move O2 to parts of submerged portions of the plant so they can get oxygen as well
- pants that grow in wet areas, either mud or partially to wholly submerged in water*
Monocot - Zea mays
- mesophyll does not have distinct layers
- conspicuous bundle sheath cells (AKA Kranz antomy) - surrounding vascular bundles, a wreath-like shape
- typical of monocots and grasses
- monocots tend to have parallel leaf veinnation
xylem
water transport in plants
bundle sheath and bulliform cells
- bundle sheath - one or more layers of parenchyma or sclerenchyma cells that surround the vascular bundles (veins) of some plants
- bulliform cells - open and close the “V” of a grass leaf, upon losing water cause the leaf to roll or fold up, large epidermal cells that occur in rows in grass leaves, and which are thought to play a role in leaf foldnig and unfolding
- turgid cells open the leaf
- folded leaf conserves water
Cuticle
- this surface blocks much of the moisture loss
aerenchyma
- tissue with large air spaces
- present on floating and emergent leaves that have blades and stalks
- helps in floatation as well as provides air passageways to the submerged roots for gas exchange
Manila hemp
- grown extensivey in Phillipines, where pineapple cloth is also made (for ceremonial purposes)
- used for tectilse, cigarette filters, “Manila” envelopes, tea bags
abscission zone
- forms across the base of the leaf stalk to facilitate leaf drop
specialized guard cells
- guard cells of eudicots and many monocots are bean shaped
- when turgid they open and close when the lose turgid
Stomatal opening
- Zeaxanthin -pigment that is able to see blue light and activates proton pumps
- Active transport - protons pumped out of guard cells, forming proton gradient
- Facilitated diffusion - potassium ions enter guard cells through voltage-activated ion channels
- Solute concentration increases - chloride ions also enter guard cells through ion channels
- guard cells become turgid - water enters guard cells by osmosis, and stoma opens
Transpiration
- the movement of water through the plant from roots, leaves, atmosphere
- leaves play critical role in transpiration
Leaf senescence
- leaf death
- individual leaves don’t have a very long life span
- leaf life spans vary by species
- in temperate species abscission is cued by shortened day length (aka cued by change in daylight)
- starches and proteins broken down into simple sugars and amino acids that are reabsorbed by the plant
- some essentional minerals also recovered
- autumn in the temperate zone yields senescing leaves containing xanthophyll and carotenoid accessory pigments
Leaf Abscission
- adaptive to winter “deserts” with little water available for photosynthesis
- leaves become a liability when they collect heavy snow causing branches and stems to snap
- ethylene (plant hormone) promotes abscission (leaf drop)
- protective layer of cork cells and suberin - leaf scar is made of this
- leaf scar is created when leaf falls*
- leaf abscission results in leaf scar
leaf scar
- protective layer of cork cells & suberin
- created when leaf falls
leaf modifications
- water storage, Aloe
- Epiphyte phyllotaxy (arrangement of leaves) stores rainwater (i.e. leaf cups to hold water, make pools of water to store its own water)
- protect floral and leaf buds
- showy
- food storage
- climbing tendrils (i.e. pea plant - allowed to climb)
Phyllotaxy
- arrangment of the leaves in a plant
leaves modified for defense
- spines are modified leaves (i.e. on a cacti)
- thorns are specialized stems (i.e. Honey locust thorn)
- prickles are extensions of epidermis or cortex (i.e. Rose prickle)
leaves trap animals
- pitcher plants
- venus fly trap
- sundew
Summary points
- leaves are arranged in distinct patterns on stems
- the epidermis provides structural support and retards water loss
- Mesophyll is the photosynthetic tissue of leaves
- xylem and phloem are the conducting tissues of leaf veins
- stomatal movements control transpiration
- autumn leaf abscission is preceded by a period of senescence
- leaf abscission is preceded by formation of an abscission zone
- some leaves are specialized for water or food storage (i.e. pitcher plant vs. plant phyllotaxy)
- leaves are modified for defense in some plants
- leaves capture animal prey in some plants