Lecture 4: Tropical forest structure and dynamics Flashcards
Key Structural characteristics of Tropical Forest Types
- Want to know what type/forest it is and what stage of succession forests are
o Because any of these forest types can be disturbed by disturbances
Secondary Forests
- Any of those types of forests (from table in previous slide) can be cleared and starting to regenerate
- Are large areas of secondary forests in tropics
- Often very close to where people are traveling
Large Increases in Secondary Degraded Tropical Forests
- Remote green areas in western Amazon = where there is primary forest left
- But most accessible areas are already secondary forests or completely cleared
- Even in largest blocks of tropical forest in world will see more secondary forest
Disturbance and succession
- Disturbance = removal of vegetation by natural or human means
o EX: Fire, windthrow - Succession = recovery of system after disturbance
o Emphasis on species composition changes
o Changes in vegetation structure
Changes in species composition and structure following disturbance
- Often have set of species that dominate after disturbance
- Completely different set of species that come in late after succession
- This results in change in forest structure over time:
o Immediately after disturbance have things growing (small statured seedlings and saplings) - 4-stage categorization:
o (1) Stand initiation stage
o (2) Stem exclusion stage
Canopy closed
Dominance by some plants
Then density dependence and exclusion
o (3) Understory reinitiation
More trees dying off
Light getting into forest
o (4) Old Growth
Trees completing lifecycle and forming gaps in forest that then are colonized
= Gap-phase dynamics
Characteristics of secondary forests
- Species that can grow rapidly (if they have lots of light)
- But tend to have much lower biodiversity than primary forests
o Small amount of tree species
o But some species that are really specialized on secondary forests
EX: Red-tailed monkeys common in secondary forests - Can also be good habitat for large mammals
o Elephants (if not hunted) - Dominance by early-successional “pioneer” species
Often easy to distinguish primary from secondary forces by canopy structure
- Smooth canopy layer in secondary forest
- Especially at stem exclusion stage
- Aerial photo: Similar views all over forest
- Recognize primary forest in the back:
o Broccoli structure with some gaps
o Lumpy
o More heterogeneous - Secondary forest more smooth
o Fewer species in primary forest
Arrested Succession
- Things can go awry
- Certain species take over and prevent further regeneration
- Most extreme example:
o In SE Asia: Non-native grass (Imperata) forms dense mats and is fire-promoting
o During dry season, these stands burn off
o Hot fire sweeps through
o Management nightmare
o Monoculture of imperata
Tropical forest gaps: signature feature of primary (old-growth) forests
- Gaps more frequent in old-growth forests
- When a tree falls in the forest creates a gap
Anatomy of a Gap
- Trees fall over because of
o Root disease
o Or wind-throw - Often tree completely uprooted
- Get tip-up mound and pit formed
o Pit contains exposed mineral soil (low nutrient status)
o Crown of tree
Most of nutrients of tree are in crown and nutrients have been displaced
Now nutrients are where the crown is may 40 m away
o Mound at root ball - Characteristic type of disturbance
- = complicated mess with particular structure
Gaps more important in tropics
- Gaps prevalent in any forest
- But more important as structural element in tropics b/c sunlight can be directly overhead
- In temperate, often have sun coming in at maximum angle (temperate)
o So unlikely to have full direct sun in middle of day - Whereas in forest, will have sunlight directly above
o = more light and stronger signal
Tip-up mound
- Formed by trees uprooting
- Results:
o Get tip-up mound w/ distinct microhabitats
o Exposure and mixing of mineral soil
o Over time have tip-up mounds forming, get pit-mound topography
= lumps and troughs through forest
Legacies of individual tree death - Secondary forests (recovered from agriculture) where there haven’t been trees in long time will be
level - But primary forests will have uneven forest floor topography
Gap phase dynamics
- In primary forest have gap-phase dynamics
o Young gaps that are bigger then get closed up
o Idealized structure could be over-simplified - Remote-sensing map
o Completely open areas down to forest floor are red
o But all gaps are heterogeneous
Characteristics of secondary and primary tropical forests
- Primary forest o Forest that’s never been logged o High Basal area Area of stem per area of ground is high Because have large trees/large stem diameters o Large seeds, animal dispersed o Have really big lianas o Epiphytes Takes time for them to be deposited o Presence of downed-woody debris o Pioneer species present maybe in gaps - Secondary forest o Forest with small trees = secondary o Small seeds o Typically have non-woody vines Have smaller lianas o Less epiphytes here o Not much downed-woody debris o Pioneers dominant in forest - General diagnostic trees
Secondary or old-growth?
- A lot of pretty small trees
- No big lianas
- Many seem like similar species
- So probably secondary
Secondary or old-growth
- Old-growth
- Huge lianas
- Big gap
- Pit and mound topography
Secondary or old-growth
- Secondary
- Cecropia
- Convergent evolution:
o Trees look like cecropia in SE Asia and Africa
Secondary or old-growth
both
Secondary or old-growth
- This is an edge
- Old-growth
What organisms need primary forest (and why?)
- Pasoh in SE Asia:
o Look at many different kinds of organisms
o Which species dependent on old-growth forests:
Fruit bats
Woodpeckers
Giant gecko - All need structures associated with big old trees
o Cavities
o Old dead trees
Applied Tropical Succession: Restoring peat swamp forests
- Restoration of peat swamp
- Characteristic peat swamp organisms
- Threats to peat swamp:
o Land-clearing for shrimp farms
Peat Swamp Forests
- Peat fires that get started
o Happen if peat swamp converted to oil palm
o Problems with air quality in Singapore
o Carbon emissions had been on par with global carbon emissions - Project:
o 1 hour north of Kuala Lumpar
o Degraded area got turned over to an NGO to try to put peat swamp forest back
Putting succession to work: Swamp forest restoration
- Need to work with local people
- Raja Musa peat swamp restoration project in Malaysia
Raja Musa Swamp Forest, Malaysia
- Gigantic area of degraded forest
- Forest peat and forest as whole = sponge as water
- One of largest rice growing places is downstream of peat swamp
- If cleared peat swamp forest wouldn’t have water available
- Important for local community
Forest Restoration Steps
- Seed collection/germination/propagation o Need native species - Closure of drainage canals o Things to bring back up water table - Trial plantings o Get rid of oil palms and get back other species - Community outreach and education - Planting day coordination
Major Challenges for Restoration
- Involve local community/public outreach and education
- Initially tried to plant some valuable timber trees that would be present in primary forest
- Didn’t work
o Not adapted to open-site conditions
o Need to find species that can deal with environment/open conditions - Need to maintain water table