hearing Flashcards
what is a sound?
- alternating waveform consisting of compression and rarefaction and separating of air molecules
- an oscillating object will cause air to become more or less dense
what equation works out wavelength
- distance between each troph
- velocity/frequency
frequency (pitch)
- hear because your brain can dissociate between different frequencies
- if not it would be a continuous noise
define amplitude (decibels)
- generally expressed as a ratio
- intensity in decibels = intensity unknown sound/intensity of standard x 10log10
- standard is mean hearing threshold
- standard mean corresponds to 10 to power of -11 movement in air molecules
what is the range of human hearing
- frequency = 20-20000Hz
- Amplitude = 0-140dB
do we have lower sensitivity to low frequencies?
- yes
- sound pressure levels adjusted to A weighting
- down-scales low frequencies to acknowledge lower sensitivity
presbycusis
- as you get older you lose auditory hair cells
- influences high frequencies
- 20 y/o have lower vol of sound to perceive it than older
having ability to listen to ten octaves of hearing
- first 2 octaves = low bass 20-80Hz
- Third and forth = upper bass 80-320Hz
- fifth-seventh = mid-range 320-2560Hz
- eighth = upper mid-range 2560-5120Hz
- ninth-tenth = treble 5120 to 20,000Hz
neurons in the ear and its frequencies
- primary afferent neurons coming from cochlea - sends auditory info to brain
- you can define frequency responsiveness of any neuron in terms of a tuning curve
- neurone responds preferentially to a frequency
- neurons respond differently to different frequencies
what is the basic anatomy of the ear
- three parts
- outer = air filled, tympanic membrane (eardrum)
- middle = air filled, ossicles
- inner = fluid filled, cochlea and vestibular system
how sound moves through ear
- tympanic membrane deflects
- middle ear bones moves and pushes oval window
- membrane in oval window moves causes cochlea fluid to move back and forth
- basilar membrane moves (contains organ of hearing)
roles of the ossicles
- middle ear acts as a lever
- ossicle bones = malleus, incus and stapes
- convert high amplitude/ low force motion at eardrum into low amplitude/high force motion at oval window
- called impedance matching
what is the stapedius reflex
- 2 muscles act on ossicles
- contraction of muscles pulls stapes away from oval window
- decreases transmission of vibrational energy to cochlea
- stapedius reflex occurs in response to very loud sound
- occur during speech
- prevents hearing damage
what happens when sound gets to cochlea
- sound vibrated through ears
- outer chamber of fluid vibrating
- 3 chambers (scale) vestibule, media, tympani
- when sound comes in it shifts column of fluid back and forth
basilar membrane
- organ of corti within it
- auditory nerve comes out of it
organ of corti in detail
- does the work
- 2 rows of hair cells inner and outer
- vestibular hair cells = physical motion
- auditory hair cells = physical motion poured by sound
- embedded in tectorial membrane
- underneath is the basilar membrane
what happens in organ of corti when sound reaches it?
- vibration of basilar membrane
- cause relative motion of tectorial membrane = activates hair cells
what type of hair cells do we have?
- one cell = kinocilium
- rest of the hair cells = stereocilia
- moving towards little hairs you get inhibition
- move towards kinocilium you get activation
what is pitch place theory?
- when basilar membrane wobbles, different parts of it will be activated at different frequencies
- closer to oval window = high frequencies
- other end = low frequencies
high frequency sound activates basilar membrane where?
proximal end
what is Fourier analysis
- the ear is one
- any waveform can be decomposed into sine waves of various frequencies
explain tuning of hearing
- far more sharp by the passive mechanics of basilar membrane alone
- amplification happening
amplification happening
- inner hair cells for sensation
- outer are for amplification
- outer contain prestin so whole cell can oscillate
- they will exaggerate sound through positive feedback cycle (active undamping)
can outer hair cells generate sound
- yes
- otoacoustic emissions are echos in response to clicks delivered to ear
- ## absense indicates problem of inner ear
what is hearing most sensitive for?
- speech
- 400-3000Hz
temporary notch
- damage depends on level and duration
- anything over 85dB is potentially damaging
- mild notch at 4kHz
what happens to sound once its past as primary afference
- sound activates many areas from cochlear to auditory cortex
- arrives in brainstem via 3 neurons
- passes through colliculus and arrives at primary auditory cortex
what is the primary auditory cortex
- responsible for your perception of sound
what is the superior olive
- in brainstem
- decifers differences in loudness and timing
what is sound localisation?
- distance (range) and bearing (azimuth and elevation)
- judge distance = high frequencies travel less well (far away is dominated by bass)
- bass travels best, sibilants worst (relative attentuation)
- echoes - reverberation
- judging direction = inter-aural timing/phase differences with left and right ear
- inter-aural volume differences
- spectral colouring - loss of high frequencies through head
what is the cocktail party effect?
- to understand conversation from an individual speaking amongst many others
- involves localisation cues and vision
what are inter-aural volume differences?
- ILD
- head provides sound shadow
- better for high frequencies (whistle nerve right ear it is much louder in right)
- bass sounds volume in both ears similar
- vowels emphasised, sibilants attenuated more far away and speech becomes more bass
inter-aural time delay
-ITD
- click from left will arrive at left ear first then right soon after
- detect ITDs at 10us (1degree)
explain the Jeffress theory
- ITD detection
- involved superior olive
- neurones in this act as coincidence detectors
- neuron splits into 5 different pathways when reaching olive (5 left 5 right)
- action potentials arrive simultaneously from both ears, MSO neurone more likely to fire
- relies on differing lengths of axons and provides a neuronal map of sound location
inter-aural phase differences
- better in low frequencies
- if sound is a continuous tone, use phase difference to localise
- not useful when wavelength is shorter than head (high frequency)
- L-R difference of 360 degree sounds same as 0 degree
does your head size matter?
- big heads = large L/R time difference - use timing differences (ITD)
- small heads = not much L/R time difference - loudness differences (ILD_
- small heads = better detecting high frequency
what is the cone of confusion?
- sounds emanating from different locations can produce identical ILD and ITD profiles at two ears
- e.g., sounds from behind can sound like the front
- if you tilt or turn head it alters the cone so a previously ambiguous sound is now localised
what does the pinna of your ear do?
- outer ear
- attenuates sounds from certain directions and amplifies from others
- invisible to low frequencies (bass)
- coming from behind it can shield high frequencies more = different spectral content
what sounds are easier to localise
- narrow bands of frequencies and gradual onsets and offsets are harder to localise
- longer, more intense broadband sounds attract attention
auditory reaction times
- typically 140-160ms
- visual reaction time is slower at 180-200ms
- frequency matters
- 1kHz = optimal reaction
startle responses
- loud sound 120dB
- evokes blink at 40ms and neck contraction at 80ms
- mediated by brainstem
- bypasses cortex via reticular formation 80ms
can a startle improve simple reaction time
- yes
- signal accompanied by loud sound reaction time is faster