Feathers & Plumages Flashcards

1
Q

Add labels to the following diagram of a feather:

A
How well did you know this?
1
Not at all
2
3
4
5
Perfectly
2
Q

What are all feather structures made of?

A

Keratin, a proteinaceous connective tissue (the same basic material of human fingernails)

How well did you know this?
1
Not at all
2
3
4
5
Perfectly
3
Q

What is a feather follicle?

A

A feather follicle is a small, epidermis-lined pit in the skin of a bird from which a feather grows and to which it remains attached.

How well did you know this?
1
Not at all
2
3
4
5
Perfectly
4
Q

What’s the difference between the pennaceous and plumulaceous portion of a feather?

A

Pennaceous describes the portion of the vane that is relatively flat and has a defined shape.

Plumulaceous describes the portion of the vane for which the loose, fluffy barbs are not structured into flat vanes.

How well did you know this?
1
Not at all
2
3
4
5
Perfectly
5
Q

In one sentence, describe how feathers are formed.

A

A feather emerges from a feather follicle like a thin-walled straw that becomes unfurled along its long axis, and its branched structure is created by the intricate fusing of fine fibres.

How well did you know this?
1
Not at all
2
3
4
5
Perfectly
6
Q

What is meant by ‘dorsal’ and ‘ventral’ feather surfaces?

A

The dorsal feather surface is the top surface of the feather while the ventral feather surface is the bottom surface of the feather.

So, for example, in flight, the dorsal surface would be facing the sky, while the ventral surface would be facing the ground.

How well did you know this?
1
Not at all
2
3
4
5
Perfectly
7
Q

What is meant by the ‘proximal’ and ‘distal’ ends of the feather?

A

The proximal end of the feather is the portion nearest to the feather follicle (bird’s body), while the distal end is distant to the feather follicle.

How well did you know this?
1
Not at all
2
3
4
5
Perfectly
8
Q

What are pin feathers?

A

Pin feathers are developing feathers that look like oddly thick, plastic rods enclosed within a bluish or greyish waxy covering called a sheath.

How well did you know this?
1
Not at all
2
3
4
5
Perfectly
9
Q

What is the function of the sheath on developing feathers (or pin feathers)?

A

The sheath protects the nascent feather as it grows. It then dries and dies, splitting and falling away (or is preened away by the bird).

How well did you know this?
1
Not at all
2
3
4
5
Perfectly
10
Q

What are fault bars and why do they sometimes develop?

A

Fault bars are transverse regions on a mature feather with slightly different colouring or texture that correspond to different stressful days of feather growth. For example, short-term periods of extreme nutritional deficiencies.

Often these fault bars are most apparent in the tail feathers.

How well did you know this?
1
Not at all
2
3
4
5
Perfectly
11
Q

True or false?

Feathers evolved from reptilian scales.

A

False.

Although feathers have some features in common with the reptilian scale, the columnar nature of the feather follicle itself is a unique feature. Thus, developmentally, from the creation of the follicle forward, all of the details of feather growth are novel and have no direct parallels to other animals.

How well did you know this?
1
Not at all
2
3
4
5
Perfectly
12
Q

Which other living creatures have feathers?

A

Birds are the only living creatures that have feathers, making them quite unique!

How well did you know this?
1
Not at all
2
3
4
5
Perfectly
13
Q

How do we know that some theropod dinosaurs had “proto-feathers”?

A

Numerous non-avian theropod fossils have been found with clear imprints of integumentary appendages surrounding the body, like pigmented halos around fossilized limbs, torsos, heads, necks, and tails.

Additionally, these appendages were found to have been made from beta-keratin, which is known only to occur in bird feathers and reptile scales.

How well did you know this?
1
Not at all
2
3
4
5
Perfectly
14
Q

Since we know most feathered dinosaurs could not fly, what was the purpose of the variety of feather forms seen across many theropod lineages?

A

It is unclear what the original function of ancient and, later, modern feathers were for dinosaurs but they seem to have diversified rapidly to serve many functions, such as protective covering, social advertising, and camouflage.

The aerodynamic or flight function of feathers only evolved later, closer in time to the evolutionary radiation of birds themselves.

How well did you know this?
1
Not at all
2
3
4
5
Perfectly
15
Q

What’s the difference between pterylae and apteria?

A

Pterylae are feather tracts—the areas of a bird’s skin to which feathers are attached—whereas apteria are regions of bare or less-feathered skin between the feather tracts.

How well did you know this?
1
Not at all
2
3
4
5
Perfectly
16
Q

What is the pterylosis of a bird?

A

The pterylosis is the detailed arrangement of feather tracts and bare patches across the body of a bird.

The pterylosis varies among groups of birds. For example, waterbirds such as penguins have a continuous pterylosis, presumably to help prevent water from penetrating to the bird’s skin and chilling it.

How well did you know this?
1
Not at all
2
3
4
5
Perfectly
17
Q

What are the two basic physical environments a bird’s feathers interact with?

A

A bird’s feathers encounter fluid media (in the form of air and water) and electromagnetic radiation (in the form of heat and light).

How well did you know this?
1
Not at all
2
3
4
5
Perfectly
18
Q

What are the SIX main groupings of feathers?

A
  1. Downy feathers: fluffy undercoat for insulation;
  2. Contour feathers: covers the torso and acts as a sunscreen, windbreaker, and raincoat;
  3. Strong pennaceous wing and tail feathers for flight;
  4. Semiplumes: insulation around the edges
  5. Ventless bristles around the mouth for feeding functions;
  6. Filoplumes: ornamental feathers for signalling to potential mates or rivals
How well did you know this?
1
Not at all
2
3
4
5
Perfectly
19
Q

What are down feathers and what function do they serve?

A

Down feathers are entirely plumulaceous, soft, and fluffy feathers that typically lack a rachis. They function as lightweight, but high-quality insulation.

Downy feathers are especially important for small birds living in cold climates, as well as for birds that float on, dive into, or swim in water.

How well did you know this?
1
Not at all
2
3
4
5
Perfectly
20
Q

What are precocial birds?

A

Precocial birds are already well-feathered with down when they hatch. They also have substantial powers of thermoregulation and locomotion, making them far more independent and able to feed themselves.

For example: the domestic chicken, many species of ducks and geese, waders, rails, and the hoatzin.

How well did you know this?
1
Not at all
2
3
4
5
Perfectly
21
Q

What are altricial birds?

A

Altricial birds are hatched relatively undeveloped and, in many cases, naked or with sparse down; such helpless young require complete parental care.

For example: Robins and most songbirds, woodpeckers, and herons**.

How well did you know this?
1
Not at all
2
3
4
5
Perfectly
22
Q

What’s the difference between natal down and body down?

A

Natal down are the feathers that cover the body of a hatchling bird, whereas body down are the feathers that lie under the contour feathers in many adult birds.

Penguins, loons, petrels, auks, geese, and ducks have these feathers for thermal insulation.

How well did you know this?
1
Not at all
2
3
4
5
Perfectly
23
Q

What bird is known for its incredibly soft and warm down feathers, which are used in the manufacture of clothing and bedding?

A

The Common Eider

Think: eider down duvets! Female ducks (and other waterfowl) actually pluck their own feathers and use them to line their nests to keep their hatchlings warm.

How well did you know this?
1
Not at all
2
3
4
5
Perfectly
24
Q

What are contour feathers?

A

Covering most of the surface of an adult bird, contour feathers give a bird its characteristic shape or outline.

The plumulaceous proximal end provides insulation while the pennaceous distal end provides a barrier against the elements (think of overlapping roof shingles).

How well did you know this?
1
Not at all
2
3
4
5
Perfectly
25
Q

What function do contour feathers serve?

A

Contour feathers form an effective barrier against sunlight, wind, and rain, while also helping to insulate and streamline the body (during flight or swimming).

How well did you know this?
1
Not at all
2
3
4
5
Perfectly
26
Q

How do most feathers manage to be effectively water repellent?

A

Water beads on the surface of most feathers due to air spaces (and trapped air) between the barbs and barbules.

Additionally, the uropygial gland at the base of their tail excretes oils that birds use to maintain and waterproof their feathers.

How well did you know this?
1
Not at all
2
3
4
5
Perfectly
27
Q

If most feathers repel water effectively, why do birds like cormorants and anhingas have to dry their wings by spreading them in the sun?

A

Cormorants and anhingas actually have feathers that are relatively easy to wet, which helps them to dive under water without the buoyant effect of trapped air bubbles. This makes them faster swimmers, but requires them to dry off by sunning themselves after feeding.

How well did you know this?
1
Not at all
2
3
4
5
Perfectly
28
Q

How do contour feathers remain organized on a bird’s body, without becoming dishevelled?

A

Tension in a set of tiny muscles attached to the contour feathers keeps them positioned correctly.

These muscles also allow the bird to move its feathers voluntarily, not individually, but in one mass movement of either raising (fluffing) or lowering (sleeking) them.

How well did you know this?
1
Not at all
2
3
4
5
Perfectly
29
Q

Give some examples of modified contour feathers.

A

The crest feathers of the Palm Cockatoo (Australia); the straw-like crown feathers of the Black-Crowned Crane (Africa); the long, elegant plumes of many herons and egrets; the fancy plumes, breast plates, and capes of many birds-of-paradise are all modified contour feathers.

How well did you know this?
1
Not at all
2
3
4
5
Perfectly
30
Q

What percentage of a bird’s total weight does its feathers typically account for?

A

Although the feathers of some birds seem to make up half of their volume, the feather coat commonly accounts for only 5-10% of a bird’s total weight.

How well did you know this?
1
Not at all
2
3
4
5
Perfectly
31
Q

What are flight feathers?

A

Flight feathers are long, strong pennaceous feathers that make up the majority of the wing and tail surfaces of birds.

How well did you know this?
1
Not at all
2
3
4
5
Perfectly
32
Q

What are remiges and rectrices?

A

Remiges are the flight feathers of the wing;

Rectrices are the flight feathers of the tail.

How well did you know this?
1
Not at all
2
3
4
5
Perfectly
33
Q

Why are flight feathers in the wing (remiges) typically asymmetrical, with one vane being slimmer than the other?

A

The leading edge of a wing feather (the vane that cuts into the air) is typically thinner than the trailing edge (or posterior vane), which is an adaptation that stabilizes the feather under the pressures of air currents during flight.

34
Q

What makes flight feathers different from most other feathers (aside from their length and shape)?

A

Flight feathers are firmly attached to the bones of the wing or tail bone, either directly or indirectly via ligaments. Other feathers are attached to the skin via the feather follicle.

35
Q

What is meant by the terms ‘primaries’ and ‘secondaries’?

A

The ‘primaries’ are the outermost, distal remiges attached to the manus bone (think of the “fingers” of large raptors).

The ‘secondaries’ are the inner proximal remiges attached to the ulna (think of the “windows” of large raptors).

36
Q

What are coverts?

A

Coverts are the smaller, overlapping feathers that overlie the bases of the flight feathers of the wing and tail.

37
Q

Many birds have feather modifications that have evolved either to exaggerate or reduce sound. What might be the functions of these traits?

A

Minimizing sound is useful for predation, as we see with owls, while exaggerating sound is associated with display behaviours and sexual selection, as is seen in many species of manakins, particularly the Club-winged Manakin.

38
Q

What are semiplumes?

A

Semiplumes are contour feathers that have lost the pennaceous portion of their vanes, as they are located in areas relatively protected from the elements by the contours of the bird’s body (e.g. in the bird’s armpit). Their function is to help to supply insulation and provide a certain amount of form as well.

39
Q

What are bristles?

A

Bristles are specialized contour feathers in which the rachis is stiffened and lacks barbs. They are functional feathers, typically located on the head, around the base of the beak, used for detection and feeding.

40
Q

What are rictal bristles?

A

Rictal bristles are stiffened feathers that project from the base of the beak, like hairs.

Many birds, particularly those that catch insects (flycatchers and nightjars) have rictal bristles for sensing their prey but also for protecting their eyes from dust and debris.

41
Q

What are filoplumes and what are they used for?

A

Filoplumes are hair-like feathers with an extremely slender, bare rachis and a tiny tuft of barbs (if any) at the tip.

Filoplumes have distinctive sensory receptors next to their follicles, which help monitor movement within the feather coat, detecting changes in feather positions that might be caused by wind or body movements.

42
Q

What is meant by the term ‘plumage’?

A

Plumage refers either to a bird’s entire feather coat, or to the set of feathers produced by a particular partial or complete molt.

43
Q

What is molting?

A

Molting is the process of shedding and replacing some or all of the feather coat with newly grown feathers.

Over time, feathers become brittle, faded, and frayed through wear and tear and exposure to the elements.

44
Q

What happens when a bird loses a feather?

A

Typically, when a single feather is lost, the feather follicle will be stimulated to produce a new one right away.

This, however, does not constitute a molt. Once or twice per year, a bird will shed and replace some or all of its older, worn feathers and replace them with fresh ones.

45
Q

How many times does a bird typically molt in its lifetime?

A

An individual bird typically experiences two series of molts in its lifetime:

  1. Those that occur as the bird matures from chick to adult, and
  2. Those that occur in annual cycles throughout the bird’s adult life.
46
Q

What can the appearance of the plumage following a molt tell us about a bird?

A

Because the appearances of the resulting plumages vary predictably within a species, they can often be used to determine an individual bird’s age.

47
Q

What is the definitive plumage of a bird?

A

The definitive plumage is the feather coat that, once attained by a bird, does not change significantly in color or pattern for the rest of its life.

In other words, its final adult plumage.

48
Q

What are subadult (or immature) plumages?

A

Subadult plumages come before the definitive plumage, and usually only differ in the coloration or patterning of the feathers.

49
Q

At what age do birds reach their definitive plumage?

A

The age at which birds reach definitive plumage varies greatly among bird species. Many songbirds take less than a year, whereas long-lived birds like large raptors, gulls, and ocean-going birds retain their subadult plumages for longer, sometimes for many years.

Example: Bald Eagles don’t attain their definitive plumage until their 4th or 5th year, while “three-year gulls”, like Ring-billed Gulls, and “four-year gulls”, like Greater Black-backed Gulls only attain adult plumage in their 3rd and 4th years respectively, with each year featuring a markedly different plumage patterning and coloration, enabling birders to estimate their age.

50
Q

Which bird is the slowest to mature and attain its definitive plumage?

A

Albatrosses may take as long as 7 or 8 years to reach their definitive plumage.

51
Q

How many molts do birds undergo in a year?

A

Some birds only undergo one complete molt each year;

Many songbirds in temperate regions molt after the breeding season and then undergo a partial molt in winter or spring;

Few birds that live in harsh habitats, where their feathers are quickly abraded and eroded (like Marsh Wrens & Bobolinks, which fly through abrasive reeds and grasses), will molt completely twice per year.

52
Q

Not all colourful breeding plumage is the result of a complete molt. Give an example of a bird whose more vivid seasonal plumage is a result of “wear-and-tear”.

A

Male Purple Finches have a dull plumage in the fall, after their annual molt in the late summer, but as the grey barbules wear away over winter, they reveal the bright cherry-red breeding plumage of the underlying barbs just in time for the following year’s breeding season!

53
Q

What is meant by ‘basic’ and ‘alternate plumage’?

A

‘Basic plumage’ is the feather coat that a bird wears for the longest period within a year, between molts and other plumage. (Example: non-breeding plumage, since this is the feather coat worn for the most amount of time each year.)

‘Alternate plumage’ is the feather coat that results from a partial molt of the basic plumage before breeding. (Example: breeding plumage, since this is the feather coat transitioned to in late winter / early spring as the bird prepares to “get jiggy with it”.)

54
Q

What is eclipse plumage and give an example of a bird that exhibits it?

A

Eclipse plumage is the set of dull-coloured feathers worn briefly after the breeding season in adult birds such as ducks. These feathers provide some protection for the males while they molt their brightly-colored feathers. It “eclipses” his usual bright plumage.

Most male ducks in eclipse plumage resemble their female counterparts. While females undergo the same prebasic molt in later summer, their appearance doesn’t change much.

55
Q

What are the four molting strategies?

A

The four molt types are basic (once per year), alternate (two molts per year), complex (‘supplemental’ plumages like juvenal variations and eclipse plumage), and, if no supplemental plumages develop, simple.

Therefore, the four molting strategies are:

  1. Simple basic (one molt per year)
  2. Complex basic (one molt each year revealing different plumages until young bird reaches adulthood)
  3. Simple alternate (two molts per year)
  4. Complex alternate (two molts with transitional plumage between them)
56
Q

How do birds lose their feathers when they molt without compromising their ability to fly and feed themselves?

A

The timing of a molt is such that a bird loses a progression of feather tracts, allowing it, for the most part, to still fly, get around, and feed.

  • Woodpeckers, who use their tails as a sturdy brace while feeding, molt the second innermost retrices first, progressing to the outermost feathers. Only once all the tail feathers have been replaced do the two strong, central retrices molt.*
  • Some waterbirds, on the other hand, like loons and grebes, drop all their remiges simultaneously, leaving them unable to fly for as long as a month during molt. But then they don’t need to fly in order to feed.*
57
Q

What is the typical pattern of wing and tail molts?

A

Wing molt often starts at the boundary between the primaries and secondaries, progressing outwards towards the distal primaries and proximal secondaries.

Tail molt proceeds outward from the central retrices.

58
Q

Explain how the rate of molt varies with latitude and the extent of seasonal change in the climate.

A

Birds generally molt when they are not engaged in other energetically demanding activities, such as feeding young, migrating, or breeding.

  • Birds breeding at polar latitudes molt rapidly because soon they must head south for the winter. The abundance of food and continual daylight for foraging in the arctic undoubtedly helps these birds in gathering the resources they need to do so.*
  • In some tropical species, on the other hand, molts may be prolonged to several months because there is no urgency to migrate and there is an abundance of resources.*
59
Q

What is preening?

A

Preening is the feather care a bird provides from the outside.

In preening, a bird grasps a feather near its base and then slides its bill along the shaft towards the tip of each feather, smoothing the barbs to re-establish any broken connections and remove stale oil and dirt.

60
Q

Explain THREE reasons birds need to preen.

A

Preening:

  1. Keeps the feathers neat and clean,
  2. Preserves their streamlining and insulating effect, and
  3. Helps remove external parasites (ectoparasites), such as flies, ticks, and lice.

Because the mature feather is a dead structure, it has no circulatory system for internal maintenance. Therefore, a bird needs to provides all feather care from the outside.

61
Q

What is the uropygial gland?

A

The uropygial gland is an oil gland located at the rump, at the base of the rectrices.

62
Q

What purpose does the uropygial gland serve?

A

The uropygial gland secretes oil that many birds spread throughout their feathers using their beaks and feet. This oil acts like a conditioner that prevents feather barbs and barbules from becoming brittle and breaking prematurely, which, in turn, both directly and indirectly, helps to keep feathers water repellent.

63
Q

Name THREE ways, other than preening, a bird can keep its feather coat clean.

A

Birds keep their feather coats clean through bathing (in water, snow, or even dust). Bathing behavior tends to be quite consistent within the avian families, with most species preferring water baths.

64
Q

How does dust bathing help to keep feathers clean?

A

Dust bathing may remove excess oils that attract ectoparasites. Alternatively, dust bathing may desiccate or clump ectoparasites already in the oils, making it easier to preen them off.

Fun fact: dust bathing is reported most often for species that spend a lot of time on the ground such as House Sparrows, Wild Turkeys, and other terrestrial birds.

65
Q

Why do birds sun themselves?

A

Suggested purposes for sunning include:

  1. Conditioning the feathers by keeping them supple through limited heating,
  2. Fast drying their feathers (cormorants and anhingas),
  3. Killing, deterring, or repositioning ectoparasites, and
  4. Saving energy by taking up solar heat through the feather coat.
66
Q

What is the behaviour known as ‘anting’?

A

Anting is something birds do to control active parasites.

In passive anting, a bird simply stations itself among a swarm of ants, permitting them to run all over its body and move in and out among the feathers.

In active anting, a bird picks up an ant or other chemically potent object (such as a millipede, plant material, or even cigarette butt) and deliberately rubs it on its feathers.

67
Q

How do birds’ eyes/vision differ to ours?

A

Birds’ eyes:

  1. Can see a greater portion of the electromagnetic spectrum than we do (slightly longer and shorter wavelengths, like UV);
  2. Have more types of color receptors than we do; and
  3. Have the anatomical equivalent of color filters that can further fine-tune how light is received by their color receptors.
68
Q

How is the color of any object (like a feather or skin) determined?

A

The color of any object is determined by what happens to light when it interacts with the object; and how this information is interpreted in the visual system of the observer.

White feathers reflect the full spectrum of long to short wavelengths of visible sunlight, and therefore appear to be white. Alternatively, when only a subset of wavelengths, like red, return to the viewer after hitting a bird’s feathers, those wavelengths stimulate specific receptors that correspond to a recognition of red.

69
Q

What are the TWO mechanisms by which light produces colors in skin and feathers?

A

Subtractive coloration: Caused by pigments (molecules, or assemblages of atoms) that absorb and interact with light at the atomic level, changing or removing light.

Additive coloration: Involves mechanisms that are influenced by the structural arrangements of materials (e.g. keratin), which reflects and transmits light in specific ways.

70
Q

What are the two main classes of pigments found in a bird’s feathers?

A

The two main classes of pigments are melanins and carotenoids.

71
Q

What effect does melanin have on the color and patterning of a bird’s feathers?

A

Color: Melanins, in general, are excellent light absorbers and mainly produce the earth tones in bird plumages, varying from the darkest blacks and subtle grays to muddy browns, red-browns, yellow-browns, and pale yellows.

Patterning: Because melanins are synthesized within each individual cell, their distribution can be very tightly controlled (cell-by-cell level control) and used to create extremely intricate patterns, such as those we see in owls, shorebirds, and pheasants.

72
Q

What effect do carotenoids have on the color and patterning of a bird’s feathers?

A

Color: Generally made of long carbon chains, carotenoids absorb and selectively re-emit light from very specific parts of the visible light spectrum, often producing bright reds, oranges, and yellows.

Pattern: Carotenoids are synthesized only by plants, algae, and fungi; therefore, birds must acquire them in their diet, either by eating plants or by eating something that has eaten those plants. As such, the color distribution is more general in nature than tightly-controlled.

73
Q

What are two other, more exotic pigments we see in some birds’ feathers?

A

Porphyrin: Complex, nitrogen-containing molecules related to haemoglobin, which are most striking in the purplish reds and greens of the turacos, a family of colorful African birds.

Psittacofulvoids: Another unusual group of pigments, as their name suggests, are found only in parrots, producing many of these birds’ vivid hues.

74
Q

What does a bird’s pigmentation signal to its potential breeding partners and rivals?

A

Because pigments are produced in the body and/or obtained through diet, they can carry information about the health and vigor of an individual bird. This can help birds advertise their overall desirability as healthy mates, or undesirability as strong competitors.

75
Q

Not all colors are produced through pigments (or pigments alone). What other key mechanism influences feather color?

A

Feathers are structurally able to scatter and refract light. This is called “structural color”.

Feathers are made of keratin which has a refractive index sufficiently different from air to cause substantial refraction. Additionally, pigments, especially melanin, have a refractive index distinctive from both air and keratin. As light travels from one medium through another, it becomes refracted, changing its color and perception.

76
Q

What color/s do incoherent scattering result in?

A

Incoherent scattering randomly scatters all wavelengths of light, leading to the appearance of white feathers (reflectance of all the colors in the spectrum cumulatively).

77
Q

What is coherent scattering and what color/s does it result in?

A

Coherent scattering occurs in feathers when the air bubbles, melanin, and keratin are arranged in very specific patterns relative to one another in ways that separate and reinforce only certain wavelengths of light. As a result, all known structural colours beyond white in birds are produced through coherent scattering.

Example: the electric-blue Plum-throated Cotinga absorbs all incoming electromagnetic radiation, while selectively scattering blue wavelengths of light.

78
Q

What is meant by ‘crypsis’?

A

Crypsis is patterning or coloration that allows a bird to remain concealed by blending into its environment. I.e. camouflage.

79
Q

What is meant by ‘disruptive coloration’?

A

Disruptive coloration is a type of camouflaging coloration in which patches, streaks, or other bold patterns help break up and distract from the overall shape of a bird.

80
Q

What is meant by ‘countershading’?

A

Countershading is a pattern of coloration in which a bird is darker on top than below, with the effect that, viewed from below, they blend into the sky and viewed from above, they blend into the ground.

This pattern is prevalent in open-country birds that are vulnerable to distant predators hunting by sight. It is also common in birds that spend much time on or flying above water, such as loons, grebes, ducks, auks, gulls, terns, and many pelagic birds.

81
Q

What is the difference between sexual dimorphism and monomorphism?

A

Sexual dimorphism is when males and females of the same species differ in aspects of their size, color, or form.

Monomorphism describes species in which males and females are similar in outward appearance.