Exam 3 Flashcards
sound
Vibrations that travel through the air or another medium as pressure waves of condensation and rarefaction, and can be heard when they reach a person’s/animal’s ear
What are the fundamental properties of sound, and how are they measured? What do we perceive them as?
- Frequency: Number of cycles per second, measured in Hertz (Hz). Perceived as pitch.
- Amplitude: Magnitude of the pressure wave, measured in decibels (dB). Perceived as loudness.
What happens to sound when it impinges on objects in the environment?
It can be…
- Reflected by the object, creating an echo
- Transmitted, with some of the sound going through the object
- Absorbed by the object
How is sound affected by our head and outer ear?
Our head’s position can influence how much sound reaches our ears, and can create a sound shadow if it blocks the sound waves from one ear. The outer ear amplifies the sound and funnels it to the eardrum.
outer ear
Includes the pinna and ear canal. This is the ear that collects sound, amplifies it, and channels it to the eardrum.
middle ear
In between the eardrum and cochlea, the middle ear helps protect the inner ear and also performs impedance matching through the use of 3 ossicles
impedance matching
More force is required to produce a given vibration in a fluid (such as in the cochlea) than in air, so the ossicles of the middle ear amplify the vibrations of the ear drum by concentrating force from a large area onto a small area (the oval window) and acting like a lever
hearing
The detection and analysis of vibrations transmitted to the ear as pressure waves in a medium such as air. A specialized form of mechanoreception.
simple sound
A pure tone, containing only 1 frequency
complex sound
A sound containing more than one frequency with different amplitudes and relative phases
spectrum of a sound
The frequency composition of a sound
harmonics
frequencies in a sound that are integer multiples of some fundamental frequency
timbre
The character of a sound (beyond its pitch and loudness), determined by the spectrum of harmonics in the sound
fundamental frequency
The frequency in a sound that is perceived as the pitch, and which all the other harmonics are multiples of
Fourier analysis
The mathematical process of breaking down a complex waveform into its component sine waves
inner ear
The part of the ear that completes transduction and spectral frequency analysis, as well as balance. The vestibular system in the inner ear works on balance. The cochlea contains the receptor cells for hearing.
What happens when pure tones of different frequencies are presented simultaneously?
They interact with each other. If they’re in-phase with each other (their waveforms line up), they sum together. If they’re out-of-phase (their waveforms are exactly opposite), they cancel each other out.
cochlea
The site at which vibrations of the stapes and inner ear fluid are transduced to neural responses. Divided into 3 fluid-filled compartments, with the scala vestibuli above the scala media, and the scala tympani below that, separated from the media by the basilar membrane, on which rests the organ of Corti.
organ of Corti
On the basilar membrane below the scala media, the organ of Corti has 3 rows of outer hair cells and 1 row of inner hair cells (the receptor cells for hearing)
hair cells
The auditory receptor cells, located on the organ of Corti. Top is covered in hair-like stereocilia, which contact the tectoral membrane and are bent according to the movement of it and the basilar membrane. Contacted by afferent projections of spinal ganglion cells from deep within the cochlea, as well as efferent nerve endings from the brain.
What are the steps of transduction in the auditory system?
- Vibrations from the stapes against the oval window of the cochlea cause upward and downward movement of the scala media
- Basilar membrane moves with the scala media, more than the tectoral plate moves, creating shearing forces on the hair cells
- Hair cells’ tip-links stretching causes K+ channels to open, allowing potassium to flow in
- Entry of K+ opens voltage-gated calcium channels
- Calcium influx triggers graded release of glutamate from vesicles at the base of the hair cell
- Ganglion cells fire action potentials (or don’t) depending on amount of glutamate received
What sound parameter is mapped throughout the auditory system?
Frequency (through the tonotopic map)
How does the cochlea accomplish frequency analysis?
The basilar membrane is stiffer at the base, and wider and looser at the apex. This causes high frequencies to cause max vibration at the base, and low frequencies to cause max vibration at the apex. This results in an internal tonotopic map of frequencies
Eustachian tube
Links the middle ear with the naso-pharyngeal cavity so that the middle ear can adjust to changes in atmospheric pressure
3 ossicles in the middle ear
- Maellus (hammer)
- Incus (anvil)
- Stapes (stirrup)
The stapes is the bone that presses against the membrane of the oval window to send vibrations into the cochlear fluid.
perilymph
The fluid in the scala tympani and scala vestibuli in the cochlea, similar to extracellular fluids