Exam 1 Flashcards
There are 6 steps of rumination. What are they, in order?
- sharp contraction of reticulum
- forces digesta and fluid to cardia
- inspiration of air against closed glottis
- creates vacuum in the thoracic esophagus
- bolus of digesta sucked into esophagus
- regurgitation of bolus + fluid of mouth
- excess liquids pressed out and swallowed
- mastication
- chewing on alternated sides of jaw with each bolus
- bolus is re-formed and re-swallowed
Describe the parotid salivary gland.
- Paired
- at the base of ear to posterior end of mandible
- most important of the salivary glands
- largest (typically >50% of total)
- highest production of saliva
- 40-50% of total saliva production
- serous (watery)
- strongly buffered
- responds to stimulation from
- Mouth, esophagus
- reticulorumen
What is a ruminant?
4-compartment (compound) stomach
Giraffids
- ruminants
- examples
- okapi
- giraffe
What occurs in the first 1/3 of gestation?
- 4 stomach compartments
- esophageal groove
- cell types (Chief, parietal) in abomasum
- distict sacs and pillars within the rumen
- omasal folds (laminae)
How does a concentrate selectors eating habits affect their anatomy?
- rapid passage, high fermentation rate
- smaller rumen
- thinner wall, fewer large pillars, evenly papillation
- smaller omasum and abomasum
- larger liver and salivary glands
- rapid rate of VFA processing needed
- shorter intestine (12-15 x body length)
- small intestine 70-73%
- large intestine 27-30%
- larger cecum +longer proximal colon
List 3 reasons why the feed efficiency (lbs of feed req. per lb of body weight gain) of a cow is inherently lower than that of a pig or chicken
- adaptations to environment
- body size
- feed quality, digestibility
How would you explain the process of rumination?
- cyclic, integrated with motility contractions
- stimulated by pressure, destension
- maximal sensitivity in cranial sac, reticulum
- destention receptors
- course material –> scratch factor
- importance of long forage diets
- maximal sensitivity in cranial sac, reticulum
Describe bulk grazers
- most recently evolved
- less selective
- consume “whole plant”
- specialize in digesting cell wall carbohydrates
- larger rumen
- longer retention times
Stratification depends on what?
- forage diets
- types of forage
- forage maturity
- processing
- moisture content
- feed intake
- mat layer dimished on high-conc. diets
- ruminant type
What are some human use of ruminants
- edible products
- meat, milk, butter, etc.
- inedible products
- fiber (wool, hair), skin/hide (leather), waste (fertilizer, fuel)
- inedible fats, tankage, endocrine extracts
- labor, recreation
- traction
- cultural (rodeo, fighting, hunting, etc.)
- other
- conservation, pest control
- financial value
What is the location and function of the nasolabial glands?
- located in the muzzle
- not true salivary glands
- secretion similar to saliva - keeps muzzle moist
- evaporative cooling (like sweat gland)
- secretes amylase
- animal licks nose, muzzle –> amylase into mouth
What is crucial in creating a milk replacer in regards to protein?
- trying to recreate casein
- protein quality
- amino acid balance and digestibility
- proteolytic enzyme development
- high pepsin, rennin
- other enzymes increase with age, exposure
- stimulating properties of casein
- curd formation, regulation of gastric emptying
What would a newborn calf’s stomach look like at birth? What would you find?
- rudimentary rumen
- thin walled
- pale appearance
- limited pailla development
- sandpaper
- highly developed, fully-functional abomasum
- about 50-70% of combined stomach volume
- preparation for milk consumption
- enzymes, HCl, volume (storage)
Tylopods
- calloused foot/camelids
- pseuruminants
- examples
- camel
- alpaca
- llama
When and where does microbial innoculation occur?
- typically within the first few hrs of life
- sources of microbes
- environment (manure, soil, etc)
- feed
- other animals
Discuss how each of these measurments correlates to the type of herbivore (bulk grazer vs. concentrate selector) and why these measurments differ between these herbivore types.
NDF, % of intake
Bulk grazers consume more NDF because they are less selective. They eat the whole plant, including the fibrous parts. Increased rumination time allows them to digest higher fiber diets
Were herbivorous dinosaurs ruminants?
- extinct prior to appearance of grasses
- ruminants first appeared ~39 million years post dino extinction
- retention time of digesta too long
- VFA broken down before absorption
- <20% energy capture
- digestive capacity issues
- 50 ton dinosaur
- 3 tons of fresh forage/day –> tons of feces/day
- no time to ruminate?
- 50 ton dinosaur
- They likely had a simple stomach–> bulk intake, low extraction approach
What is crucial in creating a milk replacer in regards to carbohydrates?
- trying to recreate lactose
- consider the enzymes present
- High lactase, low amylase and maltase (NO SUCRASE)
- Glucose, Dextrose ~ Lactose
- pre-digested lactose
- expensive
What can effect saliva output?
- eating and rumination
- nervous stimulation from the gut
- Pavlov’s response
- output can be conditioned into animals
- ex. bottle fed calves
- association of being fed w/external stimulus
- output can be conditioned into animals
- Physical nature of diet
- processing
- form and type of feed
- moisture content of feed
- more plant moisture= less saliva
Bovids
- ruminants
- example
- sheep
- bighorn
- royal antelope
- springbok
- dik dik
- cattle
- gemsbok/oryx
- goat
- bison
A) Dorsal buccal
B) Medial buccal
C, G) Ventral buccal
D, E, F) Sublingual
H) Parotid duct
I) Mandibular
J, K) Parotid
Discuss how each of these measurments correlates to the type of herbivore (bulk grazer vs. concentrate selector) and why these measurments differ between these herbivore types.
Abomasum weight
Bulk grazers have a higher abomasum weight because they are more efficient at low quality forage utilization
How does a bulk grazers eating habits affect its anatomy?
- Slower passage, fermentation (high fiber diet)
- larger subdivided rumen
- thick muscular pillars –> motility contractions
- uneven papillation –> ventral = more papilla
- larger omasum and abomasum
- smaller liver and salivary glands
- rapid VFA processing not needed
- longer intestine –> (25-30 x body length)
- small intestine 80-82%
- large intestine 18-20%
- small intestine 80-82%
What growth occurs in the last 2/3 of gestation to birth?
- general enlargement of all compartments
- initiation of rumen papilla development
What are some bulk type grazers?
- bighorn, dall, and stone sheep
- sheep
- cattle
- mouflon
Discuss how each of these measurments correlates to the type of herbivore (bulk grazer vs. concentrate selector) and why these measurments differ between these herbivore types.
Cecum weight
Concentrate selectors have a heavier cecum because their feed passes through the rumen and small intestine rather quickly and further microbe ferementation is needed at the cecum.
Pigs and Hippos
- monogastrics
- examples
- pig
- hippo
- warthog
- peccary/javelina
Excessive grain consumption can cause what disorders?
- parakeratosis (hyperkeratosis)
- keratinization of papilla
- papilla crust over “mushroom”
- increased risk of ulceration
- bacterial entry into blood stream
- ruminal and liver abscesses
- similar problem in adult animal if fed high grain diet
- associated with lactic acidosis
- keratinization of papilla
List 3 advantages that grazing/forage-fed ruminant livestock have over use of that fiber for biomass energy or cellulosic ethanol production
- a cow is a mobing bio-energy plant
- ruminants are cheaper than 1 plant
- ruminants waste is more easily disposed
Describe the sublingual salivary gland.
- paired
- located under the tongue
- smaller gland
- 10-20% size of parotid
- very low saliva production
- very high mucous
- weakly buffered
- responds to stimulation from
- mouth, espophagus (moderate)
- limited response from reticulorumen
Why are ruminants less efficient?
- body size and feed intake
- adaptation to environment
- ration quality, digestibility
- grains push 80%
- forages may push 30%
Discuss how each of these measurments correlates to the type of herbivore (bulk grazer vs. concentrate selector) and why these measurments differ between these herbivore types.
Intestine length, % of small intestine
Bulk grazers are better at nutrient absorption and must get as much out of their low quality diet as possible. A lot of absorbtion occurs in the small intestine
Antilocoaprids
- ruminants
- example
- pronghorn antelope
What are some potential replacement proteins and their downfalls?
- Animal proteins
- usually of high quality
- BSE concerns limit use
- fish protein
- generally good balance of amino acids
- relatively digestible
- prone to lipid oxidation (rancidity)
- palatibility concerns
- soybean protein (and most legumes protein sources)
- Raw soybeans
- trypsin inhibitors
- inactivation by heat
- protein allergens
- indigestible carbohydrates
- raffinose, stachyose, sucrose
- trypsin inhibitors
- Hydrolyzed soy protein
- increased digestibility, but still has above concerns
- purified soy protein (isolates, concentrates)
- removal of above concerns
- coagulation?
- limit use older calves
- Raw soybeans
Discuss how each of these measurments correlates to the type of herbivore (bulk grazer vs. concentrate selector) and why these measurments differ between these herbivore types.
CP, % of intake
Concentrate selectors pick out the highest quality parts of the plants (seeds, berries, leaves, etc. )- these habe more protein than the fibrous stems
The vagus nerve uses parasympathetic and sympathic control. Give an example of each
- Parasympathetic
- gut motility
- sympathetic
- tension receptors in stomach
How many times does an adult bulk grazer chew per day?
30,000-50,000
- This can vary with rumination type
- increased selectivity= increased eating time and decreased rumination
What are the coordinated steps of motility contractions?
- initial sharp contraction of reticulum
- same 1st contraction of rumination
- 2nd more powerful (biphasic) contraction of reticulum and reticuloruminal fold
- forces digesta past cardia into rumen
- sets off wave of contractions which pass over the whole rumen, returning to reticulum
- subsequent relaxation
- as digesta passes over ROO, small particles permitted to exit from rumen
Describe the mandibular salivary gland.
- Paired
- also called submaxillary or submandibular
- located along back of jawbone
- slightly smaller than parotid
- much lower saliva production
- high mucous
- weakly buffered
- responds to stimulation from
- mouth, esophagus
- NO RESPONSE from reticulorumen
How does a concetrate selectors jaw conformation contribute to their eating habits and mastication?
- Smaller, narrow jaw
- moderate molar surface area
- narrow muzzle
- conformation permits selectivity
- selects most digestible portion of the plant
- seeds, buds, leaves > stems
- selects most digestible portion of the plant
- slow, deliberate
- less efficient chewing, BUT
- lower fiber diet decreases need for rumination
Explain this diagram and how grasses and ruminants are related
Ruminants evolved as grasses became more available. They likely evolved so they could eat large amounts quickly and avoid predation.