Definitions Flashcards
Proximate Causation
Immediate or short-term cause of something
Example: How does the male cardinal get its red color? What is the current role of mitochondria?
Ultimate Causation
Why something exists the way it does.
Example: Why is the male cardinal red and not the female? When did mitochondria originate.
Evolution
Descent with modification.
1. Change in genetic composition of populations.
2. Cumulative changes in traits.
Requires genetic variation!
Diversification
New species
Microevolution
Generation-to-generation changes.
Example: What are the causes of evolution.
Macroevolution
Long-term changes above the species level (historical patterns)
Example: What has been the history of life on earth?
Mutation
- Error in DNA replication
- Ultimate source of all genetic variation
- CONTINUOUSLY SUPPLIES NEW ALLELES
- Single gene mutation low, genome wide high
- Arise stochastically not deterministically
Macromutations
Changes in chromosome or gene number.
Example: Deletions, duplications, translocations, inversions, fusions, point mutations.
Inversion
ABEDCF
Example: Orangutan inversion.
Fusion
Example: Human 2 fusion
Recombination
- Shuffles new genes into new combinations
- Increases variation of how genes are packaged
- Do not change in a short time scale
Example: Independent segregation, crossing-over.
Genetic Drift
Random changes in allele frequencies due to sampling error.
1. Major short-term cause of changes in allele frequencies
2. Depends on population size
3. Causes a loss in genetic variation
Example: Elephant seal, Greater prairie chicken.
Bottleneck effect

Founder effect
Founders carry unusual allele frequencies by sampling error alone.
- Form of genetic drift
- Important for some cases of speciation
Spatial subdivision
Patchy food, nesting sites or habitat
Gene flow
Movement of individuals or gametes between subpopulations.
1. Counteracts effects of genetic drift
2. Can speed up or slow down adaptive change
3. Prevents local adaptation
Example: Prevents insects from adapting to pesticide if some farmers spray. If all farmers spry then can spread favorable allele to all populations.
Hardy-Weinberg equilibrium
Allele and genotype frequencies remain constant between generations because it is a non-evolving populations characterized by:
(1) no net mutations, (2) no genetic drift (infinitely large pop), (3) no gene flow (pop isolated), (4) random mating and
(5) no natural selection)
Natural Selection
ONLY process that produces ADAPTATIVE change
Requires 3 conditions:
1. Phenotypic variation
2. Fitness differences
3. Variation in genotype
Cannot predetermine what is most useful—can only adapt to current challenges
A NON-RANDOM PROCESS
A compromise of traits that reflect historical constraints (TINKERER not engineer)
Example: Example of constraint of natural selection: Bipedalism in humans, lungs connected to stomachs in mammals, retrofitting of the testicles, laryngeal nerve in humans and giraffes, vestigal femur bone in whales and hind limb bones in snakes, mammal eye vs mollusc eye (evagination of brain vs invagination of epidermis)
Fitness
A measure of reproductive success
Directional Selection
Example: Guppie size in streams with or without predators, beak size of Soapberry bug in Flordia.
Disruptive Selection
Example: Bill size in African finches.
Stabilizing Selection
Example: Gall size of Gall fly, birth weight of human babies.
Juvenilization
Artifical selection for more juvenile like features.
Example: Wolf to dog, cattle reduced size, silver foxes breed for tameness.
Artificial Selection
Breed for a predetermined goal or path.
Example: Wild mustard cultivated into eatable kale, broccoli, cabbage, etc.
Pleiotropy
Single gene affects multiple traits.
Example: Fruit fly: bigger body size gene is same as longer egg-to-adult development time gene.
Sexual Selection
Heritable differences in ability to find mate of opposite sex.
Example: Exceptions that prove rules: Male katydids carry eggs while developing— females are much bigger. Male Phalaropes care for young, females much brighter and bigger.
Intrasexual Selection
Male-male competition (direct competition for mates).
Example: Explains behavioral/morphological traits:
(antlers, territorial behavior), post- copulatory sperm competition.
Anisogamy
Different investment per gamete.
Example: Post-zygotic care by females much higher.
Male
Produces many gametes (low investment).
Male
Produces many gametes (low investment).
Females
Produces few well-provisioned gametes (high investment).
Monogamy
One partner for entire reproductive life.
Polygyny
One male multiple females.
Example: *Sexual dimorphism (difference between sexes) is greater in polygnous species than in monogamous species.
Polyandry
One female multiple males.
Polygamy
Multiple partners for everyone.
Speciation
Populations that could once interbreed no longer can Requires:
1. Interruption of gene flow which causes isolation mechanisms.
Taxonomic species
Category of classification, generally determined by morphology.
Biological species
Species that can interbreed but are reproductively isolated from each other.
Prezygotic barriers
An isolating mechanism that prevents successful mating and fertilization.
Example: Habitat, temporal, gametic, behavioral (for species with specific mate- recognition systems), gametic
*Single gene causes mechanical isolation in land snail
Postzygotic barriers
Fertilization occurs, hybrids are inviable or infertile or break down after a few generations (can have parital barriers, one cross fertile the other isn’t).
Allopatric speciation
Caused by geographical seperations (glacier comes through, climate change, etc).
Example: Uplift of panamanian land bridge creates different lobster species.
Dispersal-mediated speciation
Pregnant lizard gets pulled out to sea and ends up on an island.
Vicariant speciation
Geographical change causes species to be seperated and change.
Example: Desert pupfish.
Ring species
Occur when ancestral population expands around a barrier. Gene flow is limited between populations. Closed ring, ends can’t breed.
Example: Western US salamader.
Sympatric speciation
Interruption of gene flow, generally in plants due to polyploidy.
Polyploidy
Increase in # of chromosomes.
Autopolyploidy
Double number of chromosomes.
Example: Potato, alfalfa, goatsbeard.
Allopolyploidy
Messed up chromosomal segregation changes number of chromosomes
More common.
Example: Cotton, tobacco, wheat, Marsh grasses (Spartina spp.)
Systematics
Establishes genealogial relationships among organisms.
Phylogeneic trees
Assembled around shared derived characteristics *have free rotation around each node.
Homology
Common ancestry means some species have same characteristics.
Example: Arms of human, cat, whale and bat.
Homoplasy
Caused by convergent evolution–traits evolved independently of each other
Example: Tasmanioan wolf tiger.
Paraphyletic
Grouping where one member is more related to some one else than the rest of the group.
Monophyletic
Grouping where everyone equally related.
Stromatolites
Earliest fossils from 3.5 bya
Phanerozoic eon
Last 550 million yrs - most fossil record comes from this eon.
Paleozoic era
550-250 may “ancient life”
Example: Biggest mass die off at brink at Premian/Triassic boundry. >50% of families and >80-90% species (due to vulcanism.
Mesozoic era
250-65 may “middle life”
Cenozoic era
65 may - present “recent life”
Example: Another big extinction at cretaceous/tertiary boundry. Dinosaurs, ammonites and rudists decimated.
Mass extinction
99% of all species are extinct.
Proof of asteroid
High concentration of iridium, shocked quartz and Chicxulub crater.
Whale evolution
Ambulocetus, rodhocetus and modern whale
Transitional species
Only identified retrospectively.
Example: Sphecomyrma freyi (ant/wasp), Archaeopteryx (dinosaur/bird)
Evolution mammals
Synapsid, therapsid, cynodont, mammals.
Example: Morganucadon, hadrocodium wui (had a derived ear structure)
Developmental biology
Reconstructy paths of change to help us understand how genetic change impacts phenotype
*tells us about homologous characteristics.
Example: Stylopod, zeugopod, autopod (Tiktaalik had neck and limb bones, Acanthostega actual tetrapod limb skeleton).
Heterochrony
Evolutionary change in the timing or rate of development.
Paedomorphosis
Retain more juvenille features.