Community Ecology (Lectures 17-23) Flashcards
what is a community?
> a group of populations coexisting in a given location.
- patterns of species diversity at a local regional scales
how do you study communities?
> scale up from two-way interactions to multi-species interactions - all the same types of interactions - predator/prey dynamics - competition - facilitation > scale down from ecosystem factors - primary productivity - climate - biogeography
how do you categorise communities?
> theoretically: set of species actively interacting
- direct competition/predation/facilitation
- indirect interactions
- gene flow
practical/ operational: group of organisms in an area smaller than a landscape that share clear physical/compositional characteristics and/or clear boundaries.
- species on a sand dune
- species in a hot spring
- species in a meadow
how do you define communities?
> closed or open systems
trophic levels
food webs
describe trophic levels for defining communities
> transfer of energy through a community
- measured by which types of species eat other types of species
- chain is kind of outdated.
describe food webs
> explore interactions among species across the trophic levels
- connectivity through predation/herbivory
- who eats who
- transfer of energy
wwhat are some issues with food webs?
> not all species fit into set categories or fit into more than one - omnivores - detritivores - parasites - pollinators > does not generally incorporate processes occurring within trophic levels - intra-specific competition - interspecific competiton
what are interaction networks?
Include: > trophic interactions - vertical links > competitive and facilitation - horizontal links - don't involve eating eachother - pairs of species interacting
what are some common ways of selecting a subse of species to study?
> taxonomic groups
- mammals, reptiles, plants
trophic guilds
- group of organisms that use the same resources
functional groups
- subset of a community that function in a similar way
types of interactions
- organisms across trophic guilds that form interaction webs through a single mechanism
- pollination, mycorrhizeal interactions, decomposition.
describe community structure
> a community has attributes that differ from those of its components
- number of species
- relative abundance of species
- nature of species interactions
- physical structure
‘fit’ of organisms to the physical environment + species interactions explain the ecological processes that control community structure
evolution equally important for understanding which set of species are found in a given community
what is species diversity?
= the number and abundance of species in a defined area
> important components of diversity
- relative abundance = proportion of individuals in a community from a single species
- species richness = the number of species in the community
- composition = the identity of the species in the community
- species evenness = tells us about the commonality and rarity of species
what is the shannon index?
> common measure of diversity
s = number of species
pi = relative abundance of each species
what is pielou’s evenness
> low evenness if one or two species dominate
> high evenness if all species have similar abundances
describe diversity and scale
> diversity will vary depending on the scale at which it is measured and the effort put in to measuring it.
species accumulation curves
higher number of samples = higher diversity
species interactions in communities
> are all species of equal importance to a community?
- Diversity?
- function?
- stability?
studies of diversity provide limited information about the processes resulting in the diversity and composition we see in given communities
describe community assembly
> understanding mechanisms of community assembly is a central goal of community ecology research
- species interactions with their environment
- the importance of the niche in understanding biological communities
- species interactions
- direct interactions
- indirect interactions
- intraspecific interactions
what are three niche theories?
> eltonian niche: the functional attributes of a species (including its trophic position).
- the environmental tolerances of a species
- a species’ relationship with food and enemies
- classic def.
grinnellian niche
- the niche of a species is determined by the habitat it lives in and behavioural adaptations to the environment
hutchinsonian niche
- N-dimensional hypervolume
- dimensions are environmental conditions and resources that determines a species’ way of life.
describe the fundamental niche constraints on community structure
> all living organisms have a range of environmental conditions under which they can successfully survive, grow, and reproduce
this range of environmental conditions conditions is not the same for all organisms
species pool - how many species are around to fill a community?
- biogeography
- evolutionary history
- ecology
describe the species pool
> same types of organisms often occupy similar environmental space
compare and contrast intra- vs. interspecific interactions
> intraspecific interactions:
- interactions among the individuals of a given species
interspecific interactions:
- interactions among individuals of a different species
the strength of these interactions can be used to understand why some species can coexist in a community and others cannot
describe direct interactions
> positive, neutral and negative interactions within or between specie that only involve the individuals (of the species) involved in the interaction
- competition for resources
- pollination
- facilitation
how do you asses interactions?
> consider direction, magnitude and stability (the net effect or long term outcome)
describe the different types of competition/facilitation
> interspecific competition -/- - resources - food, shelter etc > mutualisms +/+ - resources - services (pollination) > commensalisms +/0 - resources - habitat
how do we measure impacts of species on other species?
> relative changes in: - biomass - reproductive success - population size > models allow us to assess the effects of one species on another and then scale up to populations
what are keystone species?
> key ecological role filled by 1 species
removal of these species impacts many other species
typically not the most abundant species
impacts of removal typically disproportionate to abundance
punch above their weight
what are dominant species?
> the most common species in the community
often defined separately for each trophic level
- e.g., dominant predator, dominant producer etc
dominance is usually assumed to result from competitive exclusion
this happens in two pathways:
- transitive competition - linear hierarchy of competition
- intransitive competition - circular network of competiton
what are ecosystem engineers?
> species that create particular environmental characteristics of an ecosystem
- without them, the ecosystem changes fundamentally
are all species equally important? describe interaction strength
IS = ln((C/E)/I))
- IS = per capita interaction strength
- C = the # of target individuals in the presence of the interactor species
- E = the # of target individuals in the absence of the interactor species
- I = the # of interactor individuals
IS = (C-T)/(T*M)
- IS = per capita interaction strength
- C = mean density of target species in the presence of the interactor species
- T = mean density of a target species in the absence of the interactor species
- M = mean density of the interactor species
what is a network analysis?
> analytical approach to studying interaction webs while accounting for biological complexity.
- incorporates information on:
- strength of interactions
- direction of interactions
- commonality of interactions
common approach to the study of food webs and pollinator networks
- easiest if examining 1 type of interaction