7.7 - transport of water in the xylem Flashcards
What cells is water absorbed from, where are they located?
- root hair cells
- in the roots
Where is water transported in the plants
hollow, thick-walled tubes called xylem vessels
What is the main force that pulls water through the xylem vessels in the stem of a plant
- the evaporation of water from leaves — a process called transpiration.
- The energy for this is supplied by the sun and the process is therefore passive.
Describe movement through the stomata
- The humidity of the atmosphere is usually less than that of the air spaces next to the stomata
- As a result there is a water potential gradient from the air spaces through the stomata to the air.
- Provided the stomata are open, water vapour molecules diffuse out of the air spaces into the surrounding air.
- Water lost by diffusion from the air spaces is replaced by water evaporating from the cell walls of the surrounding mesophyll cells.
- By changing the size of the stomatal pores, plants can control their rate of transpiration.
Describe movement of water across the cells of a leaf
- Water is lost from mesophyll cells by evaporation from their cell walls to the air spaces of the leaf.
- This is replaced by water reaching the mesophyll cells from the xylem either via cell walls or via the cytoplasm.
- In the case of the cytoplasmic route the water movement occurs because:
1) mesophyll cells lose water to the air spaces by evaporation due to heat supplied by the sun
2) these cells now have a lower water potential and so water enters by osmosis from neighbouring cells
3) the loss of water from these neighbouring cells lowers their water potential
4) they, in turn, take in water from their neighbours by osmosis. - In this way, a water potential gradient is established that pulls water from the xylem, across the leaf mesophyll, and finally out into the atmosphere.
What is the main factor that is responsible for the movement of water up the xylem, from the roots to the leaves,
cohesion-tension
Describe the movement of water up the xylem
- Water evaporates from mesophyll cells due to heat from the sun leading to transpiration.
- Water molecules form hydrogen bonds between one another and hence tend to stick together. This is known as cohesion.
- Water forms a continuous, unbroken column across the mesophyll cells and down the xylem.
- As water evaporates from the mesophyll cells in the leaf into the air spaces beneath the stomata, more molecules of water are drawn up behind it as a result of this cohesion.
- A column of water is therefore pulled up the xylem as a result of transpiration. This is called the transpiration pull.
- Transpiration pull puts the xylem under tension, that is, there is a negative pressure within the xylem, hence the name cohesion— tension theory
What is some of the supporting evidence for cohesion-tension theory
- Change in the diameter of tree trunks according to the rate of transpiration. During the day, when transpiration is at its greatest, there is more tension (more negative pressure) in the xylem. This pulls the walls of the xylem vessels inwards and causes the trunk to shrink in diameter. At night, when transpiration is at its lowest, there is less tension in the xylem and so the diameter of the trunk increases.
- If a xylem vessel is broken and air enters it, the tree can no longer draw up water. This is because the continuous column of water is broken and so the water molecules can no longer stick together.
- When a xylem vessel is broken, water does not leak out, as would be the case if it were under pressure. Instead air is drawn in, which is consistent with it being under tension.
Is transpiration pull a passive process
Yes
- therefore it doesn’t use metabolic energy
Describe the features of the xylem vessel
- The vessels through which the water passes are dead and so cannot actively move the water.
- Xylem vessels have no end walls which means that xylem forms a series of continuous, unbroken tubes from root to leaves, which is essential to the cohesion-tension theory of water flow up the stem.
- Energy is nevertheless needed to drive the process of transpiration. This energy is in the form of heat that evaporates water from the leaves and it ultimately comes from the sun.
At what time of day is transpiration rate greatest? Explain your answer using information in Figure 5.
at 12.00 hours because this is when water flow is at its maximum. As transpiration creates most of the water flow they are both at a maximum at the same time.
Describe the changes in the rate of flow of water during the 24-hour period.
Rate of flow increases from a minimum at 00.00 hours to a maximum at 12.00 hours and then decreases to a minimum again at 24.00 hours.
Explain in terms of the cohesion—tension theory the changes in the rate of flow of water during the 24-hour period.
- As evaporation / transpiration from leaves increases during the morning (due to higher temperature / higher light intensity)
- it pulls water molecules through the xylem because water molecules are cohesive / stick together.
- This transpiration pull creates a negative pressure / tension.
- The greater the rate of transpiration, the greater the water flow.
- The reverse occurs as transpiration rate decreases during the afternoon and evening.
Explain the changes in the diameter of the tree trunk over the 24-hour period
- As transpiration increases up to 12.00 hours, so there is a higher tension (negative pressure) in the xylem.
- This reduces the diameter of the trunk.
- As transpiration rate decreases, from 12.00 hours to 24.00 hours, the tension in the xylem reduces and the trunk diameter increases again.
If the tree was sprayed with ammonium sulfamate, a herbicide that kills living cells, the rate of water flow would be unchanged. Explain why.
- Transpiration pull is a passive process / does not require energy.
- Xylem is non-living and so cannot provide energy.
- Although root cortex and leaf mesophyll cells are living — the movement of water across them uses passive processes, e.g. osmosis, and so continues at least for a while, even though the cells have been killed.