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Describe four processes needed for the spontaneous origin of life on Earth
The synthesis of simple organic moleculesThe assembly of these organic molecules into polymersThe formation of polymers that can self-replicate, enabling inheritancePackaging of these molecules into membranes with an internal chemistry different from their surroundings
Outline the experiments of Miller and Urey into the origin of organic compounds
• Miller and Urey recreated the conditions of pre-biotic Earth in a closed system • These conditions included a low-oxygen atmosphere, high radiation levels, high temperatures and electrical storms • Water was boiled to form vapour and then was mixed with methane, ammonia and hydrogen to form a reducing atmosphere • The mixture of gases was exposed to an electrical discharge (sparks) to simulate lightning • The mixture was then allowed to cool and after one week was found to contain some simple amino acids and complex oily hydrocarbons • Based on these findings, it was concluded that under the hypothesised conditions of pre-biotic Earth, organic molecules could be formed
State that comets may have delivered organic compounds to Earth
• Comets contain a variety of organic molecules (~20% of a comet’s tail is comprised of organic material) • Heavy bombardment about 4,000 million years ago may have delivered both organic compounds and water to early Earth
State the conditions required for the synthesis of organic molecules
Organic synthesis requires the presence of a range of inorganic molecules and an energy source to combine them into organic forms
Discuss possible locations where conditions would have allowed for the synthesis of organic compounds
• Hydrothermal vents - Regions of the ocean floor where superheated water is released from the Earth’s crust • Volcanoes - Intensive volcanic activity could provide sufficient thermal activity either on land or under the seabed • Extra-terrestrial locations - Other planets (e.g. Mars) may have been subjected to appropriate conditions (compound transfer via meteorites)
State two properties of RNA that would have allowed it to play a role in the origin of life
RNA canself-replicateRNA can act as acatalyst
Outline how RNA’s self-replicating properties would have allowed it to play a role in the origin of life
• One molecule can form a template for the production of another molecule, following the rules of complementary base pairing • Can store information in the form of a base sequence
Outline how RNA’s catalytic properties would have allowed it to play a role in the origin of life
• Could have taken the role which is now carried out by proteins (enzymes) • Modern cells use RNA catalysts (called ribozymes) to remove introns from mRNA and help synthesise new RNA molecules
Outline the theory of protobionts as preceding living cells
• Living cells may have been preceded by protobionts • Protobionts are aggregates of abiotically produced organic molecules surrounded by a membrane or membrane-like structure. • A membrane creates an internal chemical environment different to the surroundings, which could lead to optimal conditions for life activities.
Outline how prokaryotes to the creation of an oxygen-rich atmosphere
• Primordial Earth had a reducing atmosphere that contained very low levels of oxygen gas • After about ~2 billion years of prokaryotic life, a form of chlorophyll evolved in prokaryotes • This allowed for photosynthesis to occur, creating oxygen gas as a by-product via the photolysis of water
Outline the result of the creation of an oxygen-rhc atmosphere ~2 billion years ago
• The breakdown of chemicals in the atmosphere and oceans to produce oxidised compounds (e.g. CO2) • The evolution of organisms capable of performing aerobic respiration • The formation of an ozone layer which restricted UV radiation to allow for the proliferation of a wider range of life forms
Discuss the endosymbiotic theory for the origin of eukaryotes
• An endosymbiont is a cell which lives inside another cell with mutual benefit • Eukaryotic cells are believed to have evolved from aerobic prokaryotes that were engulfed by endocytosis • Mitochondria and chloroplasts are suggested to have originated by endosymbiosis
State three pieces of evidence for the endosymbiotic theory
• Mitochondria and chloroplasts have their own DNA which is naked and circular (like prokaryotes) • Mitochondria and chloroplasts have 70s ribosomes • Mitochondria and chloroplasts are roughly the same size as bacteria
Define allele frequency
The proportion of an allele within a population
Define gene pool
All the alleles present in an interbreeding population
Outline how natural selection results in a change in allele frequency in a population’s gene pool over a number of generations
• By natural selection, alleles encoding for beneficial adaptations will result in a survival advantage and lead to improved reproduction • These alleles are consequently more likely to be inherited and thus the population’s gene pool will change over generations
Discuss limitations of the definition of ‘species’
• Certain organisms (e.g. bacteria) produce asexually and therefore they do not interbreed • Some organisms are only known through fossil records, making it impossible to ascertain breeding capacity • ‘Sibling species’ look identical but don’t interbreed • Some distinct species can interbreed and produce fertile offspring • Geographically isolated organisms may never come into contact, meaning there is no information regarding their ability to interbreed
State three barriers to gene pools
Temporal isolation, behavioural isolation, mechanical isolation
Describe temporal isolation
Occurs when two species mate or flower at different timesE.G. Different frog species live in the same pond but breed at different times
Describe behavioural isolation
Occurs when two species respond to different specific courtship patternsE.G. Some crickets are morphologically identical but only respond to species specific mating songs
Describe mechanical isolation
Occurs when genital differences prevent copulation (animals) or when flowers are pollinated by different animals (plants)E.G. Galagos have distinctly shaped genitalia that will only fit other members from the same species
Explain how polyploidy can contribute to speciation
• Polyploidy is a condition in which an organism has more than two complete sets of chromosomes in all somatic cells • It is far more common in plant species as theylack separate sexes and are capable of asexual reproduction (self-pollination) • It may occur as a result of the failure of a meiotic cell to undergo cytokinesis (so chromosome replication occurs minus cell division) • Consequently gametes are diploid (2n) and resulting offspring are tetraploid (4n) • Because tetraploid offspring can no longer mate with diploid organisms(triploid offspring tend to be infertile), speciation has occurred
Outline allopatric speciation
Caused by geographical separation of populationsPopulations occupy different geographic areasE.G. Adaptive radiation of Galapagos finches
Outline sympatric speciation
Involves a reproductive, temporal or behavioural separationPopulations occupy the same geographical areasExample: Polyploidy in wheat strains
State a similarity between allopatric and sympatric speciation
Both involve the formation of a new speciesvia isolation of the gene pool from an existing species
Define speciation
The formation of new and distinct species in the course of evolution by the splitting of an existing species
Outline the process of adaptive radiation
• Adaptive radiation describes a rapid evolutionary diversification of a single ancestral lineage • It occurs when members of a single species occupy a variety of niches with different environmental selection pressures • Consequently, members evolve different morphological adaptations as a result of natural selection • Adaptive radiation results in speciation (many species from an ancestral line) and may be further enhanced by reproductive isolation • An example of adaptive radiation can be seen in the variety of beaks seen in the Galapagos finches
Outline convergent evolution
Different ancestorConverge to produce analogous structuresSpecies appearance becomes more similar over timeSpecies are unrelated (genetically different)Example: Wings in insects, birds and bats
Outline divergent evolution
Common ancestorDiverge to produce homologous structuresSpecies appearance becomes more different over timeSpecies are closely related (share genetic homology)Example: Pentadactyl limb structure in vertebrates
Outline gradualism as a pace of evolution
• Continuous change at aconstant paceover a long period of time • Arises as a result of the gradual accumulation of mutations / variations • Examples in the fossil record of gradual change with intermediate forms support this theory • An example is the evolution of the modern horse (gradual change in size and foot structure with changing habitat)
Outline punctuated equilibrium as a pace of evolution
• Evolution proceedsrapidly in burstsfor short periods of time, intermittent with long periods of stability • In periods of stability organisms become well suited to the environment, with natural selection acting to maintain characteristics • Equilibrium is punctuated by a rapid environmental change (e.g. volcanic eruption, meteor impact) which leads to directional selection • Gaps in the fossil record and the lack of intermediate forms for many species support this theory • Strata in the fossil record that show the appearance of many new species following a mass extinction support this theory
Define transient polymorphism
• A situation when there are two alleles in a gene pool (polymorphic) and one allele is gradually replacing another • This is due to a strong environmental selective pressure causing directional selection to eliminate one allele
Outline an example of transient polymorphism
Antibiotic resistance in bacteria
Define balanced polymorphism
• A situation when there are two alleles in a gene pool (polymorphic) and the frequency of the two alleles in not changing • Occurs when selective pressures promote the coexistence of the two alleles (i.e. heterozygous advantage) causing a stabilising selection
Outline an example of balanced polymorphism
• Sickle cell anaemia is controlled by a single gene mutation • Individuals homozygous for the sickle cell allele have abnormally shaped red blood cells that cause anaemia • Individuals homozygous for the normal blood cell allele are highly susceptible to malarial infection • Malaria parasite is less successful at infecting sickle-shaped blood cells • In areas where malaria is common, heterozygous individuals experience an advantage over either form of homozygote • These individuals are more likely to survive and reproduce, leading to a balance in the frequency of the two alleles (heterozygous advantage)
Outline the method of radiocarbon dating for dating rocks and fossils
• All living things are built from carbon-based organic matter • While alive, this carbon content exists as a mixture of two isotopes -12C(stable) and14C(radioactive) - maintained in constant proportions • When an organism dies, the proportion of radioactive14Cbegins to decrease as it is no longer being replaced from the environment • The proportion of14Cremaining can be used to identify the age of a sample • Carbon-14 analysis is only an effective means of dating for sample up to ~60,000 years of age as it has a half life of only 5,730 years
Outline the method of potassium-argon dating for dating rocks and fossils
• Longer range dating can be accomplished by dating the rocks around a fossil to determine an age range (i.e. relative dating) • These dating techniques can only be undertaken on igneous rocks, not on the fossils themselves or the sedimentary rock in which they are found • 40K is released in lava from active volcanos and decays into40Ar with a half life of 1,300 million years • Using the time of the volcanic eruption as a zero time point, the age of the strata in which the fossil was found can be determined • As any40Ar would have been released during the eruption as a gas, levels of40Ar in the strata provide an indication of relative age
Define half-life
The half life of a radioisotope is a measure of the time taken for the radioactivity to fall to half its original levelIn other words, it is a measure of the time taken for half the atoms in a radioactive sample to decay
Describe the major anatomical features that define humans as primates
• Forward facing eyes, giving stereoscopic vision • Large brain volume relative to body • Opposable thumbs capable of grip
Outline the trends illustrated by the fossils of Ardipithecusradius, Australopithecus including A. afarensis and A. africanus, and Homo including H. habilis, H. erectus, H. neanderthalensis and H. sapiens
fckn do it
State that, at various stages in hominid evolution, several species may have coexisted
Several species of hominid may have co-existed at the same time: • Homo habilismay have co-existed with various species ofAustralopithicus • Homo neanderthalensislikely co-existed withHomo sapiens
Discuss the incompleteness of the fossil record
• Fossilisation is an exceptionally rare occurrence that requires an unusual combination of special conditions • Most living things tend to decompose rapidly (or be scavenged) following death • Fossilisation tends to favour hard body parts (bone, teeth, shells, etc.) and exposed fossils will soon be weathered / destroyed • Only a small percentage of fossils have been discovered - fossilisation favours species that were long-lived and widespread
Discuss uncertainties about human evolution resulting from the incompleteness of the fossil record
• Individual fossils may not be representative of species (e.g.Homo floresiensis- ‘Hobbit’ man) • Very few complete skeletons have been discovered, and so paleoanthropology is an inductive (data-poor) science • Many conclusions have been drawn on limited data and are frequently re-interpreted in the light of new discoveries
Discuss the correlation between the change in diet and increase in brain size during hominid evolution
• A change in habitat in Africa ~2.5 million years ago may have prompted the emergence of Homo species from forest to savannah • This necessitated a change of diet to include meat (more protein), which increased the skull capacity and brain size of hominids • Improved diet quality provided energy to support greater brain function and learning capacity (positive correlation) • Activities resulting from improved cognition (such as group hunting and cooking food) enabled hominids to eat a wider variety of food